The Human Nervous System is the
Most Sensitive to Environmental Toxins
By
Kaye H. Kilburn
from Approach to a Consensus for
the Basis of Regulation of Environmental Chemicals, (eds) J. D.
Arcos, et al. Special Issue of Environmental Carcinogenesis Reviews
Part C of Journal of Environmental Sciences and Health, Marcel
Dekker, Inc. (1991)
Every school child is taught that the human nervous
system evolved to apprehend and perceive the environment. It registers,
and discriminates with great sensitivity, light, color, sound,
vibration, touch, pain, position, odor, and taste. Could this
capacity be it's Achilles heel? Sensitivity may be its undoing.
We have accepted the fact that other organs are
highly sensitive to environmental toxins, such as the lung to
asbestos and Isocyanates, the liver to carbon tetrachloride and
vinly chloride, the kidney to cadmium and phenacetic, and the
skin to methacrylate and urushiol. However, intuition and history
favor the brain and nervous system for sensitivity. If this is
true, why has it escaped our attention until recently?
Perception of damage to the brain and nervous system
may be masked by its wonderful redundancy. Such effective damage
control is carried out by substituting function and redirecting
impulses so that until reserves are nearly gone, only a slowing
of response may be seen. And because slowing is regarded as an
ordinary manifestation of aging, the damage is concealed. Even
considerable slowing may be overlooked in clinical neurological
evaluations, which are only qualitative and at best measure one
subject's nervous system against that of another, making fine
estimates of time impossible.
Quantative measurements focused on speed appear
to be the key to the appraisal of many subtle changes in function,
elusive until recently. Just as spirometry measures the speed
of expiratory airflow from the lung, reaction time, speed of the
sway-movement, and blink speed (for example) measure the time
needed to perform diverse functions. Ideal comparisons would be
to follow up an individual across time or across an episode of
exposure, which may be brief or prolonged. Short of such ideal,
but very rare cases, comparisons between populations exposed to
an agent or situation or to a site, such as a workplace or neighborhood,
require controls.
In a world increasingly
polluted, finding a group of human subjects to compare to may
be a most difficult problem.
Although the origins of neurotoxicology are lost
in prehistory, systematic observations of palsy and colic from
lead were made by Benjamin Franklin and of erethism (irritation),
tremor, and cachexia in mirror silverers exposed to mercury in
amalgams by Ramazzini and by Kussmaul, and immortalized in "hatters
shakes" and madness by Lewis Carroll. Remarkably, despite
this long history, the focusing on neurotoxins and their targets
in the brain has been belated, and there has been an apparent
lack of a unifying body of concepts concerning their detection
and mechanisms of action.
As a first step toward rectifying these deficiencies,
I suggest that we reexamine the significance of the fact that
the human nervous system, with its special sensory endings, is
a highly evolved apparatus for apprehending and perceiving the
environment. However, this sensitivity may be its very undoing.
The intuitive hypothesis has been advanced that among the systems
of living organisms, the nervous system is, in fact, the most
liable to damage from environmental toxins.
Damage may be masked
against detection, however, because subtle
dysfunction is often concealed
by the nervous system's remarkable redundancy and ability to substitute
function, or it is simply overlooked in clinical evaluations which
are usually only quantative. Unless various functions are appraised
quantitatively, dysfunction may be attributed to anxiety, tension,
aging or a deficient endowment of intelligence.
The Critical keys are quantitative measurements
of diverse functions which can be compared to past performance,
to predicted scores, or to the behavior of unexposed individuals
or populations. The menu of the variety of measurements spreads
subtly from the affective and symptomatic, through cognitive functions
which require the subject's cooperation such as recall and fund
of information, to the measurement of unconscious or automatic
functions which have previously been in the sphere of the neurophysiologist.
Symptoms of altered affective states or mood are
common in every day experience, particularly fatigue, headache,
irritability and depression. The application of quantitative methods
such as the Minnesota multi-phasic personality inventory (MMPI)
or the profile of mood states (POMS) to the appraisal of affective
states and the correlation of these with the outcome of psychological
tests for cognitive functions, such as perceptual motor speed,
sensory motor reaction time, verbal and visual-spatial memory
and design analysis, have begun to provide insight into their
relations with cognitive and neurological functions.
Based on profound and prolonged neurobehavioral
impairment in individuals such as on the battlefield or as prisoners
of war or other ordeals, "post traumatic stress" has
been advanced as an explanation for dysfunction. This thesis has
also been offered to explain the effects of occupational or environmental
exposure to neurotoxins such as organophosphate pesticide over-dose
or exposure to trichloroethylene in well water. Although some
affected subjects display anger or appear outraged, more often
they appear apathetic or depressed. To argue that affective disorders
derange neurobeyhavioral function begs the question, mistakes
an outcome for a cause and ignores the mind-body unity which is
remarkably well demonstrated in the integration of the human nervous
system.
Although the human mind has many facets, the integrative
position argues that affective disorders which we characterize
as confusion, tension-anxiety, depression, anger, fatigue and
absence or vigor in the profile of mood states, arise form the
apprehension by the damaged but still perceptive brain of its
disordered function. Thus, affective disorders are the outcome
of toxic encephalopathy and not its imitators by adversely affecting
testing. Such damage appears analogous to the situation after
massive localized brain damage from vascular accidents (strokes),
when all these affective states may ensue together, and depression
and/or confusion are frequently obvious on even casual observation.
Tests originally developed to study alcoholism,
to select recruits for pilot training, to explore the effects
of aging, and to estimate educable children for school placement,
are used together with other tests that have been developed to
measure toxicity of materials at work, for assessing effects of
environmental toxins on the human nervous system.
It would be remiss to proceed further without crediting
David Wechsler's contributions in devising two standard instruments.
One is an immediate learning (recall and memory) scale and the
other is an adult intelligence scale(1,2).
Wechsler cautioned that "the information obtained from intelligence
tests is relevant to the extent that establishes and reflects
whatever it is one finds in overall capacity for intelligent behavior."
The tests used to measure intelligence served primarily as a means
to an end. "In this respect, the tests do not differ essentially
from devices employed by physicists to measure heat by the use
of thermometers or thermoelectric couples."
Wechsler combined 11 tests in his WAIS(2)
and standardized each. The verbal scale included information,
digit span, vocabulary, arithmetic, comprehension, and similarities.
The performance scale included picture completion, picture arrangement,
block design, object assembly, and digit symbol. Essentially,
these were pencil and paper tests suited for literate and language-aware
individuals. Others, such as Cattell, deliberately avoided language
and literacy in devising "Culture Fair," which consists
of four groups of designs: logical series, maximal difference,
design completing, and definition establishment and application,
for estimating human intelligence(3).
The application of tests to environmental or occupational
groups should follow Wechsler's suggestions that the abilities
measured broadly characterize human behavior, and that intelligence
is multi-phasic and multi-determined with a judgment of overall
competency, and finally, that test scores and competency in coping
effectively with the environment are not necessarily closely related.
Testing the conscious level, nevertheless, requires cooperation,
is strongly dependent on understanding language and on capacity
to handle numbers, and is also somewhat sensitive to attitude
and feeling states.
Attempts to reduce the influence of intelligence,
vigilance, cooperation, and feeling states on the final integrative
assessment, and to encompass other CNS processes, led to the measurement
of automatic neurophysiological functions. Examples include: Nerve
conduction, the blink reflex, body balance, visual and auditory
evoked potentials, and peripheral vibration and temperature senses.
Traditionally these measurements have been restricted to specialized
laboratories and have required such large investments of time
per subject that they have seldom been employed for epidemiological
or even clinical studies. Their use has been limited to confirmation
rather than detection. This is a pity, because there is evidence
that balance is reversibly affected by ethanol(4),
that blink, particularly the initial motor response, is reduced
by trichloroethylene(5), that nerve conduction
is reduced by lead(6), and that the visual
evoked potential, the responses to flash versus pattern are significantly
changed in aluminum intoxication(7).
All of these methods broaden the clinical neurologists'
qualitative estimates with quantitative ones for detecting subtle
differences across time or delays in function induced by exposures.
Investigator certainty or "comfort" is greatest when
the automatic or neurophysiological tests show abnormality or
dysfunction in concert with the ones requiring subject cooperation,
and that these also keep track with affective appraisal scores
and symptoms.
Clearly, the first application of testing for the
affecting-symptomatic area, the conscious-cooperative-cognitive
area and the subconscious-automatic neuro-physiological one, is
to understand a single patient and relate his impairment or dysfunction
to toxic exposures.
Application beyond this subject to many individuals
in a group is an epidemiological survey. Sensitivity, i.e., the
ability to detect effects, is frequently greater in an epidemiological
survey because small differences when compared to suitable populations
may be significant while the individual must be looked at in terms
of 95% confidence intervals or deviations from his measured or
estimated prior level of function.
One among the critical issues is how to estimate
previous levels of function when they are unavailable. For some
rests, years of school completed is a sensitive predictor; in
others, age is a major determinant. Balance exemplifies a function
with a U-shaped curve with age, showing improvement during growth
and development, a plateau during adulthood and progressive impairment
in advanced age. Poor balance also relates to frailty and occurs
typically in the eighth or ninth decade of life(8).
Subjects exposed to either occupational or environmental chemicals
can be studied using these methods.
Workers breathe and contact chemicals and metals
while employed for perhaps 40 hours a week. "Environmental
neighbors" usually receive diluted doses down-stream or down-wind
from industrial processes or their dump sites but for up to 168
hours per week. Although studies or painters with exposure to
lead and solvents and rayon workers exposed to carbon disulfide
have produced quantitative data to explore these problems, it
is now clear that chemicals in dumps, chemical exposure during
hazardous material clean-up particularly with the use of ingenious
disposal methods such as incineration and injection into exhausted
petroleum wells, offer new challenges involving mixtures of chemicals.
Chemical waste may be defined functionally,
as mixtures too expensive to refine, provided they can be conveniently
"lost." Thus, the toxic clean-up workers, the
HAZMAT teams, the well-head injectors and perhaps firemen, above
all others, are at greatest risk.
It is instructive to consider
this century's lessons from lead poisoning - which was manifested
by colic and palsy in the last century - that learning deficits
and behavioral problems occur in young children in whom increased
lead absorption is without a threshold(9,10). In fact, dentine
lead, a marker of cumulative lead dose is correlated with increased
incidence of "problem behavior" in children compared
to controls (who themselves consist of subjects with greatly increased
body burdens as compared to pre-industrial populations). Consequent
to lead poisoning in young children, a constellation of neurobehavioral
disorders including hyperactivity has been called minimal brain
dysfunction (MBD) or attention deficit disorder (ADD). It has
been suggested that in these children, the monoaminergic systems
are dysfunctional, that myelin synthesis and repair are defective,
and that nervous system development is impaired. In any case,
the neurotoxicity of lead appears to be irreversible(10).
Recently, exposure to high impact noise from artillery
and rifle fire, well known to cause deafness, was shown in Finnish
soldiers to also adversely affect balance(11).
Our preliminary studies on ironworkers, men who assemble structural
steel in notoriously noisy construction sites using heavy machinery
including air impact hand tools, show hearing loss and impaired
balance which are highly correlated (personal observation). Lead
toxicity has also been associated with hearing loss(12)
as has solvent exposure, illustrating that both chemical, and
physical agents can adversely affect cochlear and vestibular functions
of the eighth cranial nerve, and the brain which governs them.
Neurobehavioral testing of a population chronically
exposed to trichloroethelene (TCE), trichloroethane, chromium
and other chemicals in their well water for up to 25 years showed
impairment in most domains including memory (recall), perceptual-motor
speed, dexterity, balance, blink, and in recognizing logical series,
differences, design completion and applying definitions in designs.
Comparison to a national reference group showed that these impairments
were statistically significant(13). Comparison
to a similar population in the same state uncovered another well
water contamination incident with TCE and other chlorinated solvents.
At Woburn (MA) a cluster of leukemia cases stimulated investigations
which demonstrated high prevalence of fatigue, headache, memory
loss, and difficulty of concentrating(14).
Specific demonstrating of prolonged R, of blink reflex response
was attributed to trigeminal nerve poisoning by TCE(15).
Individual patients have shown neurobehavioral impairment
from a brief, subfatal exposure to hydrogen sulfide, carbon monoxide
or PCBs, and from prolonged occupational exposures to formaldehyde,
styrene and epoxy resin systems (including ethyl and methyl aniline)
used to make prosthetic limbs, and to mixtures of toxic chemicals
injected into abandoned oil wells. Exposures ranged from single
elements such as lead and aluminum - and simple compounds such
as hydrogen sulfide, carbon monoxide and formaldehyde - to complex
mixtures such as were found in the Livingston (LA) train derailment
of cars containing 21 different chemicals to multiple industrial
solvents in the ground water in Woburn (MA) and Tucson (AZ).
Firemen exposed to pyrolysis products of PCBs in
a single dose superimposed upon years of repeated subclinical
exposures, showed neurotoxicity months after the incident(15). Perhaps comparison of firemen in the PCB
incident to firemen not in the incident concealed measurable dysfunction
which would be obvious if these "control" firemen were
compared to men without exposure. Quantitative expressions of
neuropsychological function enhance the opportunity for finding
associations with body burdens of such materials(16).
Recently, alcohol was once again identified as an
added-load exposure which, for example, reinforced lead-induced
cognitive dysfunction and added to workers' irrationality and
behavioral problems(17). It appears that
the effects of neurotoxicity may be unrecognized in an exposed
population until an episode stimulates investigations which lead
to demonstration of impairment.
Much remains to be done in
this field. Despite its long history, some branches on its developmental
tree are puny or missing. It would help to standardize the descriptive
language, and not only group the tests into logical categories,
but to standardize them so as to increase the comparability between
the work of various investigators on the same toxic substance
or of similar occupationally or environmentally exposed populations.
What is also needed, is a
clear understanding of the relationships between measures of affective
disorder (MMPI or POMS), symptom inventories, cognitive or perceptual
testing which requires cooperation, and testing of subconscious
and automatic (neurophysiological) performance. As the apparatus
for these latter tests is made robust for field use, progress
should accelerate. Comparisons across these axes should reveal
interrelationships and map the dysfunctions of the nervous system.
Quantitative comparisons should increase specificity and predictive
capacity. They would improve epidemiology by providing measurements
linking it to biochemical and biological markers(18). Rapid progress may
be imperative in order to conserve neurobehavioral competence
in the population. The field must mature far behind its substantial
but somewhat incoordinate beginnings, just described.
The ultimate Hogartian picture shows that - as human
discriminatory functions fall -
not only will it
be impossible to find unexposed subjects for comparison, but society's
resolve and ability to identify the problems and solve them will
ebb.
The examples of alcohol, heroin, cocaine, marijuana,
"crack," and "speed" which alter perception
temporarily are disquieting. Insidious permanent loss is more
so. Perhaps it is more than a historical speculation that the
fall of the Roman Empire was linked to lead, ingested after it
was leached from containers by wine and water.
Contributing to this nightmarish
scenario is evidence that the effects of neurotixins mimic aging
such that the population's brains age faster than other organs
to produce, at worst, epidemic Alzheimer's disease. The
economic and political implications are awesome. The realization
that the effects of solvents are analogous to aging has led to
an elegant method of analysis. It consists of constructing regression
equations for age and specific neurophysiological or neuropsychological
functions for populations exposed to potential neurotoxins and
for control groups. Larger are coefficients in those exposed suggest
cumulative effects from extended periods of exposure and confirm
the "form frustra" of the nightmare.