http://www.collembola.org/taxa/katiidae.htm - Last updated on 2009.09.29 by Frans Janssens
Checklist of the Collembola: Katiannidae

Familia Katiannidae Börner, 1913, sensu Bretfeld G, 1999:13 key
Genus Number
of species
Arborianna1
Betschurinus1
Dalianus1
Gen.nov.1
Gisinianus1
Katianna50
Katiannellina1
Katiannina2
Keratopygos1
Millsurus1
Neokatianna1
Parakatianna12
Polykatianna11
Pseudokatianna14
Rusekianna4
Sminthurinus89
Stenognathellus4
Vesicephalus7
Zebulonia1
Katiannidae203
Genera with number of species
Sminthurinus bimaculatus from France
2005 © Losacco, S.
Sminthurinus elegans from the USA
2006 © McClarin, J.


Genus Gen.nov. Greenslade P in Ardron, PA, 2009:16
Genus nov. sp. nov. from the UK
Third antennal segment with papilla and long fine setae
2009.04.01 © Ardron, P.
Det. Greenslade, P.

Species sp.nov. Greenslade P, 2009:il
Genus nov. sp. nov. from the UK
After Ardron PA, 2009 p.16
Genus nov. sp. nov. from the UK
Mucrones
2009.05.20 © Janssens, F.
Det. Greenslade, P.
Genus nov. sp. nov. from the UK
Female sub anal appendage
2009.05.20 © Janssens, F.
Det. Greenslade, P.

  Forma instar.1 Greenslade P, 2009:il
Genus nov. sp. nov. from the UK, 1st instar
2009.02.23 © Barton, T.

  Forma instar.2 Greenslade P, 2009:il
Genus nov. sp. nov. from the UK, juvenile
2007.03.18 © Cornwall, N.J.
Genus nov. sp. nov. from the UK
2009.02.23 © Barton, T.
Genus nov. sp. nov. from the UK
2009.02.23 © Barton, T.
Genus nov. sp. nov. from the UK
2009.03.06 © Barton, T.
Genus nov. sp. nov. from the UK
2009.04.07 © Shaw, P.
Genus nov. sp. nov. from the UK
2009.04.21 © Ardron, P.
Genus nov. sp. nov. from the UK
2009.04.21 © Ardron, P.

  Forma instar.3 Greenslade P, 2009:il
Genus nov. sp. nov. from the UK, juvenile
2007.03.18 © Cornwall, N.J.
Genus nov. sp. nov. from the UK, juvenile
2007.03.04 © Cornwall, N.J.
Genus nov. sp. nov. from the UK, juvenile
2007.03.18 © Cornwall, N.J.
Genus nov. sp. nov. from the UK, juvenile
2007.03.18 © Cornwall, N.J.
Genus nov. sp. nov. from the UK
2009.02.23 © Barton, T.
Genus nov. sp. nov. from the UK
2009.02.23 © Barton, T.
Genus nov. sp. nov. from the UK
2009.02.23 © Barton, T.
Genus nov. sp. nov. from the UK
2009.02.23 © Barton, T.
Genus nov. sp. nov. from the UK
2009.03.06 © Barton, T.
Genus nov. sp. nov. from the UK
2009.03.06 © Barton, T.
Genus nov. sp. nov. from the UK
2009.03.06 © Barton, T.
Genus nov. sp. nov. from the UK
2009.03.06 © Barton, T.
Genus nov. sp. nov. from the UK
2009.03.15 © Barton, T.
Genus nov. sp. nov. from the UK
2009.04.21 © Ardron, P.

  Forma instar.4 Greenslade P, 2009:il
Genus nov. sp. nov. from the UK, juvenile
2007.03.04 © Cornwall, N.J.
Genus nov. sp. nov. from the UK, juvenile
2007.03.18 © Cornwall, N.J.
Genus nov. sp. nov. from the UK, juvenile
2007.03.18 © Cornwall, N.J.
Genus nov. sp. nov. from the UK, juvenile
2007.03.18 © Cornwall, N.J.
Genus nov. sp. nov. from the UK, juvenile
2007.03.18 © Cornwall, N.J.
Genus nov. sp. nov. from the UK
2009.02.23 © Barton, T.
Genus nov. sp. nov. from the UK
2009.03.06 © Barton, T.
Genus nov. sp. nov. from the UK
2009.03.06 © Barton, T.
Genus nov. sp. nov. from the UK
2009.03.06 © Barton, T.
Genus nov. sp. nov. from the UK
2009.03.06 © Barton, T.
Genus nov. sp. nov. from the UK
2009.03.06 © Barton, T.
Genus nov. sp. nov. from the UK
2009.03.06 © Barton, T.
Genus nov. sp. nov. from the UK
Sub-anal appendage
2009.03.27 © Ardron, P.
Genus nov. sp. nov. from the UK
2009.04.01 © Ardron, P.
Genus nov. sp. nov. from the UK
2009.04.01 © Ardron, P.
Genus nov. sp. nov. from the UK
2009.04.01 © Ardron, P.
Genus nov. sp. nov. from the UK
2009.04.01 © Ardron, P.
Genus nov. sp. nov. from the UK
2009.04.01 © Ardron, P.
Genus nov. sp. nov. from the UK
2009.04.01 © Ardron, P.
Genus nov. sp. nov. from the UK
2009.04.21 © Ardron, P.
Genus nov. sp. nov. from the UK
2009.04.21 © Ardron, P.
Genus nov. sp. nov. from the UK
2009.04.21 © Ardron, P.
Genus nov. sp. nov. from the UK
2009.04.21 © Ardron, P.
Genus nov. sp. nov. from the UK
2009.04.21 © Ardron, P.

  Forma instar.5 Greenslade P, 2009:il
Genus nov. sp. nov. from the UK
2007.03.18 © Cornwall, N.J.
Genus nov. sp. nov. from the UK, juvenile
2007.03.20 © Cornwall, N.J.
Genus nov. sp. nov. from the UK
2007.03.20 © Cornwall, N.J.
Genus nov. sp. nov. from the UK
2009.02.23 © Barton, T.


Genus Keratopygos Christiansen, KA et Pike, E, 2002:180, -
Species megalos Christiansen, KA et Pike, E, 2002:181, -, t.t.

Genus Arborianna Bretfeld, 2002
Species cupiubae Bretfeld, 2002, t.t.

Genus Parakatianna Womersley, 1932
Species albirubrafrons Salmon, JT, 1943:6
Species anomala Womersley, 1933
Species cortica Salmon, JT, 1943:8
Parakatianna cortica from New Zealand
After Salmon JT, 1943 Pl.1 Fig.9
Parakatianna cf. cortica from the UK
2008.01.20 © Spears, G.

Species diversitata Salmon, JT, 1943:7
Parakatianna diversitata from New Zealand
After Salmon JT, 1943 Pl.1 Fig.4

Species hexagona Salmon, JT, 1941:414
Parakatianna hexagona from New Zealand
After Salmon JT, 1941 Pl.71 Fig.510

Species homerica Salmon, 1946

Parakatianna albirubrafrons
subsp. niveanoa from New Zealand
After Salmon, J.T. 1943 Pl.1 Fig.5.
Species niveanota Salmon, JT, 1943:7
Species prospina (Salmon, 1946), t.t.
Species salmoni Wise, 1964
Species spinata Womersley, 1932, t.t.
Species superba Salmon, 1946
Species viridis Salmon, JT, 1943:8
Parakatianna diversitata
subsp. viridis from New Zealand
After Salmon JT, 1943 Pl.1 Fig.11


Genus Katianna Börner, C, 1906:182
Katianna sp. from the USA
2006 © Bernard, E.
Katianna schotti
Katianna australis from the UK
2008.12.09 © Ardron, P.A.
New Katianna spp. from the UK
2009.06.06 © Ardron, P.A.
Katianna sp. from Australia
2009.08.29 © Hort, J.

Species sp.1 Delamare Deboutteville, C et Massoud, Z in Delamare Deboutteville, C et Rapoport, E, 1963:241
Species sp.2 Greenslade, P et Ireson, JE, 1986:282
Katianna sp. from Australia
After Greenslade P & Ireson JE, 1986 Fig.2

Species sp.3 Janssens F et Ardron PA in Ardron, PA, 2009:15
Katianna sp. from the UK
2006 © Barton, T.
Katianna sp. from the UK
2006 © Barton, T.
Katianna sp. from the UK
2007 © Barton, T.
Katianna sp. from the UK
2007.03.04 © Cornwall, N.J.
Katianna sp. from the UK
2007.03.04 © Cornwall, N.J.
Katianna sp. from the UK
2007.03.04 © Cornwall, N.J.
Katianna sp. from the UK
2007.03.18 © Cornwall, N.J.
Katianna sp. from the UK
2007.03.18 © Cornwall, N.J.
Katianna sp. from the UK
2007.03.18 © Cornwall, N.J.
Katianna sp. from the UK
2007.03.18 © Cornwall, N.J.
Katianna sp. from the UK
2007.03.18 © Cornwall, N.J.
Katianna sp. from the UK
2007.03.18 © Cornwall, N.J.
Katianna sp. from the UK
2007.12.05 © Barton, T.
Katianna sp. from the UK
2007.12.05 © Barton, T.
Katianna sp. from the UK
2007.12.10 © Barton, T.
Katianna sp. from the UK
2007.12.16 © Barton, T.
Katianna sp. from the UK
2007.12.16 © Barton, T.
Katianna sp. from the UK
2008.08.16 © Ardron, P.A.
Spines on head
Katianna sp. from the UK
2008.08.16 © Ardron, P.A.
Katianna sp. from the UK
2008.08.16 © Ardron, P.A.
Katianna sp.3 from the UK
After Ardron PA, 2009 p.15
Katianna sp. from the UK
2009.04.21 © Ardron, P.A.
Katianna sp. from Tasmania
2009.08.16 © Daley, A.

  Forma flammea Ardron PA, 2009:16
Katianna sp. f. flammea from the UK
2009.06.06 © Ardron, P.A.
Katianna sp. f. flammea from the UK
2009.06.06 © Ardron, P.A.
Katianna sp. f. flammea from the UK
2009.06.06 © Ardron, P.A.
Katianna sp. f. flammea from the UK
2009.07.24 © Barton, T.

Species sp.4 Janssens F et Ardron PA in Ardron, PA, 2009:16
Katianna sp.4 from the UK
After Ardron PA, 2009 p.16
Katianna sp. from the UK
2009.04.27 © Ardron, P.A.
Katianna sp. from the UK
2009.04.27 © Ardron, P.A.
Katianna sp. from the UK
2009.04.27 © Ardron, P.A.
Female subanal appendage
Katianna sp. from the UK
2009.06.06 © Ardron, P.A.
Katianna sp. from the UK
2009.06.06 © Ardron, P.A.

Species sp.5 Ardron, PA, 2009:19
Unknown sp.1 from the UK
After Ardron PA, 2009 p.19

Species sp.6 Ardron, PA, 2009:19
Katianna s.l. sp. from the UK
2008.mm.dd © Ardron, P.
Katianna s.l. sp. from the UK
2008.mm.dd © Ardron, P.
Sminthurinus nr elegans var. dorsalis from the UK
After Ardron PA, 2009 p.18
Unknown sp.2 from the UK
After Ardron PA, 2009 p.19

Species antennalis Rapoport, 1962
Species antennapartitia Salmon, JT, 1941:412
Species aurantiaca (Womersley, H, 1935:218) Greenslade, P in Houston, WWK, 1994:124
Species australis Womersley, 1932
Katianna australis from Australia
After Womersley H, 1939 Pl.1 Fig.5.
Katianna australis from Australia
After Greenslade P & Ireson JE, 1986 Fig.6
Katianna australis from the UK
After Ardron PA, 2009 p.15
Katianna australis from the UK
2009.02.01 © Ardron, P.A.
Katianna australis from the UK
2009.02.01 © Ardron, P.A.
Katianna australis from the UK
2009.02.01 © Ardron, P.A.

Species banzarei Salmon, JT, 1964:314
Katianna banzarei from Antarctica
After Davis, G. in Greenslade, P., 2006 Fig.81.

Species cardoni (Delamare Deboutteville & Massoud, 1963:237) Rapoport, 1968
Species cobold Börner, 1907
Species coeruleocephala Handschin, 1920
Species decora Börner, 1907
Species drummondi (Womersley, 1932) Greenslade P in Houston, 1994:125
Species gloriosa (Womersley, H, 1937:157) Salmon, 1946
Arrhopalites gloriosa from Australia
After Womersley H, 1937 Fig.1m

Species guttulata Börner, 1907
Species houssayi Delamare Deboutteville & Massoud, 1963:225
Species jeanneli Delamare Deboutteville & Massoud, 1963:222
Species kerguelenensis Denis, 1947
Remark by Deharveng, L. 1981 (cfr Greenslade, P., 2007.10.09:in litt.): This species probably belongs in the genus Pseudokatianna because it lacks the generic character of spines on the head according to the figures.

Remark by Greenslade, P. 2007.10.09:in litt.: Taking into consideration the form of the mucro, the chaetotaxy of the dens, the slight antennal annulation and other features, the species probably is not a Pseudokatianna but may belong in Polykatianna, especially as Denis noted it had conspicuous interocular bosses. However, neither of these two genera are well defined, so the correct generic placement for this species is not possible to determine at present.

Species kuscheli Delamare Deboutteville & Massoud, 1963:228
Species liliputana (Nicolet, H, 1847:360) Stach, J, 1956:239, i.s.
Remark by Ellis, W.N. & Bellinger, P.F. 1973:28: Nicolet, 1847 described new species from Chile. These are also described by Nicolet in Gay, 1851[lapsus for Nicolet in Gay, 1849]. But in the first paper Nicolet cites as author of the species Nicol. in Gay. Evidently the publications appeared in a reversed order.

Smynthurus(sic) liliputanus from Chile
After Nicolet H in Gay C, 1854 Lam.1 Fig.9.

Species mnemosyne Börner, C, 1906:182, t.t.
Species mucina Womersley, 1933
Katianna mucina from Australia
After Greenslade P & Ireson JE, 1986 Fig.3

Species nunezi (Najt, 1967) Betsch, 1977
Species obscura (Womersley, H, 1936:485) Greenslade, P & Ireson, JE, 1986:275
Parakatianna obscura from Australia
After Womersley H, 1936 Fig.7a
Parakatianna obscura from Australia
After Womersley H, 1936 Fig.7b
Parakatianna obscura from Australia
After Womersley H, 1936 Fig.7c
Katianna sp. cf. obscura from Australia
After Greenslade P & Ireson JE, 1986 Fig.5

Species oceanica Schött, 1917
Species ornata Womersley, 1932
Species perplexa Salmon, JT, 1944:176
Species pescotti Womersley, H, 1935:218
Species poivrei (Delamare Deboutteville & Massoud, 1963:233) Rapoport, 1968
Species puella (Denis, 1933) Betsch, 1977
Species purpuravirida Salmon, JT, 1941:413
Katianna purpuravirida from New Zealand
After Salmon JT, 1941 Pl.71 Fig.502

Species reducta Salmon, JT, 1944:176
Species richardsi Najt, 1967
Species ruberoculata Salmon, JT, 1944:175
Species schotti Womersley, 1933
Katianna oceanica var. schoetti(sic) from Australia
After Greenslade P & Ireson JE, 1986 Fig.4
Katianna schotti from the UK
2008.04.30 © Ardron, P.A.
Katianna schotti from the UK
2008.04.30 © Ardron, P.A.
Katianna schotti from the UK, mucrones
2008.04.30 © Ardron, P.A.
Katianna schotti from the UK
Dorsal macrosetae
2008.04.30 © Ardron, P.A.
Katianna schotti from the UK
2008.05.06 © Ardron, P.A.
Katianna schotti from the UK
Peglike papilla on 3rd antennal segment
2008.05.06 © Ardron, P.A.
Katianna schotti from the UK
Spinelike setae on head
2008.05.06 © Ardron, P.A.
Katianna schotti from the UK
2008.05.06 © Ardron, P.A.
Katianna schotti from the UK
2008.05.06 © Ardron, P.A.
Katianna schotti from the UK
2008.05.06 © Ardron, P.A.
Katianna schotti from the UK
Foot complex with tenent hairs
2008.05.06 © Ardron, P.A.
Katianna schotti from the UK
2008.12.09 © Ardron, P.A.
Katianna schotti from the UK
2008.12.11 © Ardron, P.A.
 
Katianna schotti from the UK
2008.12.19 © Ardron, P.A.
 
Katianna schotti from the UK
2008.12.19 © Ardron, P.A.
Katianna schotti from the UK
After Ardron PA, 2009 p.14
Katianna schotti from the UK
2009.05.06 © Ardron, P.A.
 
Katianna schotti from the UK
2009.05.06 © Ardron, P.A.
Katianna schotti from the UK
2009.05.06 © Ardron, P.A.
Katianna schotti from the UK
2009.05.06 © Ardron, P.A.
Katianna schotti from the UK
2009.06.06 © Ardron, P.A.
Katianna schotti from the UK
2009.06.06 © Ardron, P.A.
Katianna schotti from the UK
2009.06.06 © Ardron, P.A.
Katianna schotti from the UK
2009.06.06 © Ardron, P.A.
Katianna schotti from the UK
2009.06.06 © Ardron, P.A.

Species serrae Najt, 1967
Species serrata (Schäffer, 1897) Stach, 1956
Species serrata Womersley, 1936, nec Schäffer, 1897
Species steparia (Najt, 1967) Mari Mutt & Bellinger, 1990:140
Species tillyardi Womersley, 1932
Species uschuaiensis (Schäffer, 1897) Stach, 1956
Species viretorum Najt, 1967
Species viridis Delamare Deboutteville & Massoud, 1963:237
Species viridula Womersley, 1932
Species willincki (Delamare Deboutteville & Massoud, 1963) Betsch, 1980
Species wygodzinskyi Delamare Deboutteville & Massoud, 1963:228
Katianna nr wygodzinskyi from Peru
2009.01-03.dd © Contreras Valdez, A.

Species zebra (Womersley, H, 1942:30) Greenslade, P in Houston, WWK, 1994:126
Parakatianna zebra from Australia
After Womersley H, 1942 Fig.7A
Parakatianna zebra from Australia
After Womersley H, 1942 Fig.7B


Genus Polykatianna Salmon, 1946
Species aurea (Womersley, 1932) Bruzzese, 1980
Species cremea Salmon, JT, 1949:55
Species davidi (Tillyard, 1920) Greenslade P & Wise, 1986
Parakatianna litorea
subsp. luteaterga from New Zealand
After Salmon, J.T., 1943 Pl.1 Fig.6.
Polykatianna davidi from Antarctica
After Davis, G. in Greenslade, P., 2006 Fig.78.

Species flammea Salmon, 1946, t.t.
Species intermedia (Snider, 1978) Mendoza Arviso, S, Villalobos, FJ, Ruíz Montoya, L & Castro, AER, 1999:90
Species montana (Womersley, H, 1936:483) Greenslade P in Houston, 1994:128
Parakatianna montana from Australia
After Womersley H, 1936 Fig.6a

Species polygonia Snider, 1978
Species pulchella (Womersley, 1933) Greenslade P in Houston, 1994:128
Species quinseta Salmon, 1954
Species radicula (Maynard, EA, 1951:224)
Sminthurinus radiculus from the USA
After Maynard EA, 1951 Plate XXIII Fig.378

  Subspecies pallens Maynard, EA, 1951:225
Sminthurinus radiculus f. pallens from the USA
After Maynard EA, 1951 Plate XXIII Fig.380

  Subspecies pictus Maynard, EA, 1951:224
Sminthurinus radiculus f. pictus from the USA
After Maynard EA, 1951 Plate XXIII Fig.379

Species utahensis (Snider, RJ, 1988:1)
Sminthurinus (Polykatianna) utahensis from the USA
After Snider RJ, 1988 Fig.1
Sminthurinus (Polykatianna) utahensis from the USA
After Snider RJ, 1988 Fig.2


Genus Sminthurinus Börner, C, 1901:99
Remark by ICZN. 2007:759: Placed on the Official List of Names in Zoology. Opinion 1027.

Species sp.1 Zeppelini, D, Bellini, BC, Creao-Duarte, AJ et Hernández, MIM, 2009:1166
Species sp.2 Zeppelini, D, Bellini, BC, Creao-Duarte, AJ et Hernández, MIM, 2009:1166
Species albiceps (Schött, 1896) Stach, 1956, s.i.
Species albipes (Schött, 1896) Stach, 1956, i.s.
Species alpinus Gisin, 1953
  Subspecies bisetosus Ellis, 1976
Species antennalis (Rapoport, 1962) Delamare Deboutteville & Massoud, 1963:213
Species atrapallidus Snider, 1978
Sminthurinus atrapallidus from the USA
2006.02.25 © Schimming, L.
Sminthurinus atrapallidus from the USA
2009.02.07 © Justis, S.
Sminthurinus atrapallidus from the USA
2009.02.07 © Justis, S.
Sminthurinus atrapallidus from the USA
2009.02.07 © Justis, S.
Sminthurinus atrapallidus from the USA
2009.02.07 © Justis, S.
Sminthurinus atrapallidus from the USA
2009.02.07 © Justis, S.
Sminthurinus atrapallidus from the USA
2009.04.08 © Zungri, D.

Species aureus (Lubbock, 1862) Börner, C, 1901:103, B
Smynthurus(sic) aureus from the UK
After Lubbock J, 1873 Pl.7
Sminthurinus aureus from the UK
Ventral subapical dental setae absent
2004 © Hopkin, S.P.
Sminthurinus aureus
2006 © Boldsystems.org
Sminthurinus aureus from the Netherlands
2009.01.26 © Heemskerk, R.

  Subspecies purpureus Salmon, JT, 1941:410
  Varietas alba Krausbauer, 1902
  Varietas maculata Krausbauer, 1902
  Varietas ochropus Reuter in Schött H, 1893:32
  Varietas punctata Krausbauer, 1902
  Varietas pustulata Krausbauer, 1902
Species aureussetosus Bretfeld, 2002
Species australis Najt, 1973
Species bimaculatus Axelson, 1902
Sminthurinus bimaculatus from France
2005 © Losacco, S.
Sminthurinus bimaculatus from France
2005 © Losacco, S.
Sminthurinus bimaculatus from the UK
2008.06.06 © Barton, T.

Species bisetosus Ellis, 1976
Species bullai Najt, 1973
Species cantonensis Rusek, 1971
Species castagninoi Najt, 1969
Species cingula Bagnall, 1932, s.d.
Species cingulatus Bagnall, 1921
Species conchyliatus Snider, 1978
Species concolor (Meinert, 1896) Stach, 1956
Sminthurinus concolor from the UK
Two setae on retinaculum
2004 © Hopkin, S.P.
Sminthurinus concolor from Luxembourg
After Stomp N & Weiner WM, 2005 Fig.5

Species discordipes Salmon, JT, 1949:49
Species dispar Babenko, AB, 2007:375
Species distinctus Kang J-S et Lee B-H in Kang, J-S, Lee, B-H et Park, K-H, 2005:240
Sminthurinus distinctus from Korea
After Kang J-S, Lee B-H & Park K-H, 2005 Fig.4A

Species domesticus Gisin, 1963
Sminthurinus domesticus from the UK
Two subapical ventral dental setae
2004 © Hopkin, S.P.
Sminthurinus domesticus from the UK, juvenile
2007.09.29 © Valentine., B
Sminthurinus domesticus from the UK
2007.10.13 © Valentine., B
Sminthurinus domesticus from the UK
2007.10.13 © Valentine., B
Sminthurinus domesticus from the UK
2007.10.18 © Valentine., B
Sminthurinus domesticus from the UK
2007.10.24 © Valentine., B
Sminthurinus domesticus from the UK
2007.10.25 © Valentine., B
Sminthurinus domesticus from the UK
2007.10.25 © Valentine., B
Sminthurinus domesticus from the UK
2007.11.15 © Valentine., B
Sminthurinus domesticus from the UK
2008.08.27 © Valentine., B
Sminthurinus domesticus from the UK
2008.10.16 © Valentine., B
Sminthurinus domesticus from the UK
2008.10.16 © Valentine., B
Sminthurinus domesticus from the UK
2008.11.18 © Valentine., B
Sminthurinus domesticus from the UK
2008.11.18 © Valentine., B
Sminthurinus domesticus from the UK
2008.12.05 © Valentine., B
Sminthurinus domesticus from the UK
2008.12.05 © Valentine., B
Sminthurinus domesticus from the UK
After Ardron PA, 2009 p.18

Species duplicatus Salmon, JT, 1941:411
Species elegans (Fitch, 1862) Folsom, 1928, B
Sminthurus quadrilineatus from Sweden
After Schött H, 1893 Taf.I Fig.18
Sminthurinus elegans from the USA
After Maynard EA, 1951 Plate XXIV Fig.424
Sminthurinus elegans from the USA
After Maynard EA, 1951 Plate XXVIII Fig.520
Sminthurinus elegans from the USA
After Maynard EA, 1951 Plate XXVIII Fig.521
Sminthurinus elegans f. cancellus from the USA
After Maynard EA, 1951 Plate XXVIII Fig.522
Sminthurinus elegans f. cancellus from the USA
After Maynard EA, 1951 Plate XXVIII Fig.523
Sminthurinus similitortus from the USA
After Maynard EA, 1951 Plate XXIV Fig.413
 
Sminthurinus elegans from Australia
After Greenslade P & Ireson JE, 1986 Fig.2
Sminthurinus elegans
Rare form with lateral white spots
2004 © Baquero, E.
Sminthurinus elegans from the UK
Female sub-anal appendages
2004 © Hopkin, S.P.
Sminthurinus elegans from the USA
2006 © McClarin, J.
 
Sminthurinus elegans from Sweden
2006 © Fjellberg, A.
Sminthurinus elegans from the USA
2008.05.16 © Thombs, D.
Sminthurinus elegans from the UK
2008.12.09 © Valentine., B
Sminthurinus elegans from the UK
2008.12.09 © Valentine., B
Sminthurinus elegans from the UK
2008.12.09 © Valentine., B
Sminthurinus elegans from the USA
2009.02.14 © Justis, S.
Sminthurinus elegans from the USA
2009.02.14 © Justis, S.
Sminthurinus elegans from the USA
2009.02.14 © Justis, S.
Sminthurinus elegans from the USA
2009.02.14 © Justis, S.
Sminthurinus elegans from the USA
2009.05.21 © Cowen, R.

  Varietas dorsalis Axelson, 1905
  Varietas ornata Krausbauer, 1902
  Varietas signatus Krausbauer, 1898
  Forma ochropus American authors, nec Reuter
Sminthurinus elegans f. ochropus from the USA
After Maynard EA, 1951 Plate XXVIII Fig.524
Sminthurinus elegans f. ochropus from the USA
After Maynard EA, 1951 Plate XXVIII Fig.525
Sminthurinus near aureus from the UK
After Ardron PA, 2009 p.19
 
Sminthurinus elegans f. ochropus from the USA
2009.02.08 © Justis, S.
Sminthurinus elegans f. ochropus from the USA
2009.02.08 © Justis, S.

Species exiguus (Nicolet, H, 1847:360)
Remark by Ellis, W.N. & Bellinger, P.F. 1973:28: Nicolet, 1847 described new species from Chile. These are also described by Nicolet in Gay, 1851[lapsus for Nicolet in Gay, 1849]. But in the first paper Nicolet cites as author of the species Nicol. in Gay. Evidently the publications appeared in a reversed order.

Species gamae Gisin, 1963
Species gisini da Gama, 1965

Sminthurinus glaucus from New Zealand
After Salmon, J.T. 1943 Pl.1 Fig.8.
Species glaucus Salmon, JT, 1943:2

Species gloriosus (Womersley, 1937) Greenslade P in Houston, 1994:131
Species granulatus Salmon, 1946
Species granulosus Enderlein, G, 1908:409
Species handschini Folsom, 1934
Species henshawi (Folsom, 1896) Stach, 1956
Sminthurinus (Sminthurinus) henshawi from the USA
After Snider RJ in Christiansen KA & Bellinger PF, 1998 Fig.914A
Sminthurinus henshawi from the USA
2006 © McClarin, J.

  Forma aureus Folsom, 1898
Sminthurinus aureus from the USA
After Maynard EA, 1951 Plate XXIII Fig.410
Sminthurinus henshawi f. aureus from the USA
2009.03.10 © Cowen, R.
Sminthurinus henshawi f. aureus from the USA
2009.03.10 © Cowen, R.
Sminthurinus henshawi f. aureus from the USA
2009.03.10 © Cowen, R.

  Forma megoculatus Maynard EA, 1951:233
Sminthurinus megoculatus from the USA
After Maynard EA, 1951 Plate XXIX Fig.542
Sminthurinus megoculatus from the USA
After Maynard EA, 1951 Plate XXIX Fig.544
Sminthurinus henshawi f. megoculatus from the USA
2007.03.26 © Reed, D.E.
Sminthurinus henshawi f. megoculatus from the USA
2008.01.26 © Montgomery, G.
Sminthurinus henshawi f. megoculatus from the USA
2008.01.26 © Montgomery, G.

  Forma similitortus Maynard EA, 1951:232
Sminthurinus similitortus from the USA
After Maynard EA, 1951 Plate XXIV Fig.414
Sminthurinus similitortus from the USA
After Maynard EA, 1951 Plate XXIX Fig.537
Sminthurinus similitortus from the USA
After Maynard EA, 1951 Plate XXIX Fig.538
Sminthurinus henshawi f. similitortus from the USA
2009.02.05 © Justis, S.
Det. Snider, R.J.
Sminthurinus henshawi f. similitortus from the USA
2009.02.05 © Justis, S.
Det. Snider, R.J.
Sminthurinus henshawi f. similitortus from the USA
2009.02.07 © Justis, S.
Det. Snider, R.J.
Sminthurinus henshawi f. similitortus from the USA
2009.02.07 © Justis, S.
Det. Snider, R.J.
Sminthurinus henshawi f. similitortus from the USA
2009.02.07 © Justis, S.
Det. Snider, R.J.
Sminthurinus henshawi f. similitortus from the USA
2009.02.07 © Justis, S.
Det. Snider, R.J.
Sminthurinus henshawi f. similitortus from the USA
2009.04.08 © Justis, S.

Species hygrophilus Bretfeld, 2000
Species igniceps (Reuter, 1878) Börner, C, 1901:102
Sminthurus igniceps from Sweden
After Schött H, 1893 Taf.II Fig.17
Sminthurinus igniceps from France
2007.03.17 © Lebeaux, P.

Species inexcussus Najt & Rapoport, 1965
Species insularis Najt, 1973
Species jonesi Wise, KAJ, 1970:203
Sminthurinus jonesi from Antarctica
2009.03.20 © Key, R.S.
Sminthurinus jonesi from Antarctica
2009.03.20 © Key, R.S.

Species kaha Christiansen, K et Bellinger, P, 1992:349
Species kerguelensis Salmon, JT, 1964:314
Species krausbaueri Börner, 1901, t.t.
Species latimaculosus Maynard, EA, 1951:231
Sminthurinus latimaculosus from the USA
After Maynard EA, 1951 Plate XXIII Fig.389
Sminthurinus latimaculosus from the USA
After Maynard EA, 1951 Plate XXIII Fig.390

Species lawrencei Gisin, 1963
Species lichenatus Salmon, JT, 1943:3
Species lineatus Bretfeld in Bretfeld, Poliakov & Broza, 2000
Species maculosus Snider, 1978
Species mime Börner in Voeltzkow, 1907
Species minutus (MacGillivray, 1894) Maynard, EA, 1951:234
Sminthurinus minutus from the USA
After Maynard EA, 1951 Plate XXIII Fig.402
Sminthurinus minutus from the USA
After Maynard EA, 1951 Plate XXVIII Fig.535
Sminthurinus minutus from the USA
2006.12.02 © Murray, T.
Sminthurinus minutus from the USA
2007.01.09 © Montgomery, G.
Sminthurinus minutus from the USA
2008.02.09 © Montgomery, G.
Sminthurinus minutus from the USA
2009.04.11 © Zungri, D.
Sminthurinus minutus from the USA
2009.04.11 © Zungri, D.

Species modestus Yosii, 1970
Species molinai Arlé, 1940
Species mountfordi (Womersley, H, 1939:242) Greenslade P in Houston, 1994:132
Species muscophilus Salmon, 1946
Species niger (Lubbock, 1867) Börner, C, 1901:101, t.t., B
Smynthurus(sic) niger from the UK
After Lubbock J, 1873 Pl.6
Sminthurinus niger from France
2005 © Locasso, S.
Sminthurinus niger from the UK
2006.11.01 © Brocklehurst, K.
Sminthurinus niger from the UK
2006.11.01 © Brocklehurst, K.
Sminthurinus niger from the UK
2006.11.01 © Brocklehurst, K.
Sminthurinus niger from the UK
2007.03.04 © Cornwall, N.J.
Sminthurinus niger from the UK
2007.03.18 © Cornwall, N.J.
Sminthurinus niger from the UK
2007.03.18 © Cornwall, N.J.
Sminthurinus niger from the UK
2007.12.22 © Barton, T.
Sminthurinus niger from the UK
2007.12.22 © Barton, T.
Sminthurinus niger from the UK
2008.01.10 © Valentine., B
Sminthurinus cf. niger from the USA
2008.01.21 © Moorehead, C.
Sminthurinus niger from the UK, female
2008.04.08 © Valentine., B
Sminthurinus niger from the UK, male
2008.04.08 © Valentine., B
Sminthurinus niger from the UK
2008.11.28 © Valentine., B
Sminthurinus niger from the UK
2008.12.25 © Corley, P.
Sminthurinus niger from the UK
2009.01.04 © Corley, P.
Sminthurinus niger from the UK
2009.03.10 © Valentine., B
Sminthurinus niger from the UK
2009.03.10 © Valentine., B
Sminthurinus niger from the UK
2009.04.12 © Barton, T.

  Varietas var. Stach, 1956
  Forma alpina Gisin, 1953
  Forma ochracea Axelson, 1902
Species nigrafuscus Salmon, JT, 1941:409
Species nigrescens Schött, 1917
Species nunezi Rapoport in Delamare Deboutteville & Massoud, 1963:210
Sminthurinus nunezi from Peru
2009.01-03.dd © Contreras Valdez, A.

Species obscurus Salmon, JT, 1944:173
Species oculatus Schött, 1917
Species operosus Najt & Rapoport, 1965
Species orientalis Stach, J, 1964:21
Species packardi Denis, 1933
Species pallescens Yosii, 1970
Species pallidus Womersley, 1931
Species patagonicus Najt, 1971
Species pekinensis Stach, J, 1964:22
Species planasiensis Dallai, 1969
Species procerasetus Salmon, 1946
Species pullus Bretfeld in Bretfeld, Poliakov & Broza, 2000
Species quadratus Bretfeld in Bretfeld, Poliakov & Broza, 2000
Species quadrimaculatus (Ryder, 1878) Handschin, 1928
Sminthurinus quadrimaculatus from the USA
After Maynard EA, 1951 Plate XXIII Fig.393
Sminthurinus quadrimaculatus from the USA
After Maynard EA, 1951 Plate XXIII Fig.394
Sminthurinus quadrimaculatus from the USA
After Maynard EA, 1951 Plate XXVI Fig.489
Sminthurinus brunneus from the USA
After Maynard EA, 1951 Plate XXIII Fig.400
Sminthurinus brunneus from the USA
After Maynard EA, 1951 Plate XXIII Fig.401
Sminthurinus downsi from the USA
After Maynard EA, 1951 Plate XXII Fig.377
Sminthurinus quadrimaculatus f. latipictus from the USA
After Maynard EA, 1951 Plate XXIII Fig.391
Sminthurinus quadrimaculatus f. latipictus from the USA
After Maynard EA, 1951 Plate XXIII Fig.392
Sminthurinus quadrimaculatus from the USA
Pale form
2006 © McClarin, J.
Sminthurinus quadrimaculatus from the USA
2007 © McClarin, J.
 
Sminthurinus quadrimaculatus from the USA
First instar
2007.04.12 © Murray, T.
Sminthurinus quadrimaculatus from the USA
Second instar
2007.04.15 © Murray, T.
Sminthurinus quadrimaculatus from the USA
Second instar
2007.04.15 © Murray, T.
Sminthurinus quadrimaculatus from the USA
2007.10.23 © McClarin, J.
Sminthurinus quadrimaculatus from the USA
2008.10.04 © Murray, T.

  Forma bimaculatus Maynard, EA, 1951:230
Sminthurinus quadrimaculatus f. bimaculatus from the USA
After Maynard EA, 1951 Plate XXVI Fig.490
Sminthurinus quadrimaculatus f. bimaculatus from the USA
After Maynard EA, 1951 Plate XXVI Fig.491
Sminthurinus quadrimaculatus from the USA
Form with only 2 spots
2003 © Schoenherr, J.
Sminthurinus quadrimaculatus from Canada
Form with only 2 spots
2007.05.17 © Pilon, C.
Sminthurinus quadrimaculatus from Canada
Form with only 2 spots
2007.05.17 © Pilon, C.
Sminthurinus quadrimaculatus from the USA
Form with only 2 spots
2007.12.16 © Moorehead, C.
Sminthurinus quadrimaculatus from the USA
Form with only 2 spots
2008.06.03 © Schoenherr, J.
Sminthurinus quadrimaculatus from the USA
Form with only 2 spots
2008.06.04 © Schoenherr, J.
Sminthurinus quadrimaculatus from the USA
Form with only 2 spots
2008.09.08 © Murray, T.

  Forma bipunctatus Maynard, EA, 1951:235
Sminthurinus minutus f. bipunctatus from the USA
After Maynard EA, 1951 Plate XXIII Fig.403
Sminthurinus minutus f. bipunctatus from the USA
After Maynard EA, 1951 Plate XXVIII Fig.536
Sminthurinus quadrimaculatus from the USA
Form with only 2 spots
2008.02.07 © Gross, J.
Sminthurinus quadrimaculatus from the USA
Form with only 2 spots
2008.02.07 © Gross, J.
Sminthurinus quadrimaculatus from the USA
Form with only 2 spots
2008.02.07 © Gross, J.

Species reticulatus Cassagnau, P, 1964:481
Sminthurinus reticulatus from Sweden
After Fjellberg A, 2006 ©
Sminthurinus reticulatus from the UK
2006.11.01 © Brocklehurst, K.
Sminthurinus reticulatus from the UK
2006.11.01 © Brocklehurst, K.
Sminthurinus reticulatus from the UK
2006.11.01 © Brocklehurst, K.
Sminthurinus reticulatus from the UK
2006.11.01 © Brocklehurst, K.
Sminthurinus reticulatus from the UK
2006.11.01 © Brocklehurst, K.
Sminthurinus reticulatus from the UK
2006.11.01 © Brocklehurst, K.
Sminthurinus reticulatus from the UK
2006.11.01 © Brocklehurst, K.
Sminthurinus reticulatus from the UK
2006.11.01 © Brocklehurst, K.
Sminthurinus reticulatus from the UK
2006.11.01 © Brocklehurst, K.
Sminthurinus reticulatus from France
2007.09.08 © Lebeaux, P.
 
Sminthurinus reticulatus from France
2008.03.24 © Lebeaux, P.
Sminthurinus reticulatus from France
2008.03.24 © Lebeaux, P.
Sminthurinus reticulatus from France
2008.12.07 © Jacquemin, G.

Species salti Delamare Deboutteville, 1953
Species sayi Denis, 1933
Species signifer (Fitch, 1862)
Species speciosus Yosii, 1970
Species splendidus (Womersley, 1933) Greenslade, P in Houston, WWK, 1994:133
Parakatianna splendida from Australia
After Womersley H, 1939 Pl.1 Fig.4.

Species subalpinus Itoh, 2000
Species suborientalis Stach, J, 1965:367
Species timorensis Yoshii & Suhardjono, 1992

Sminthurinus transversalis from Finnland, 1902
Det. Axelson, W.M.
2006 © Fjellberg, A.
Species transversalis Axelson, WM, 1905:794

Species tricolor Schött, 1917
Species trinotatus Axelson, WM, 1905:794, B
Sminthurinus trinotatus from Germany
2006.12.26 © Kohl, F.
Sminthurinus trinotatus from Germany
2006.12.29 © Kohl, F.
Sminthurinus trinotatus from Belgium
2008.06.09 © Vuijlsteke, M.
Sminthurinus trinotatus from the UK
2008.07.22 © Shaw, P.
Sminthurinus trinotatus juv. from France
2008.12.07 © Jacquemin, G.
Sminthurinus trinotatus juv. from France
2008.12.07 © Jacquemin, G.
Sminthurinus trinotatus from France
2008.12.07 © Jacquemin, G.
Sminthurinus trinotatus from France
2008.12.07 © Jacquemin, G.
Sminthurinus trinotatus from France
2008.12.07 © Jacquemin, G.
Sminthurinus trinotatus from the UK
After Ardron PA, 2009 p.17
Sminthurinus trinotatus from the UK
2009.09.01 © Thomason, R.
Sminthurinus trinotatus from the UK
2009.09.01 © Thomason, R.

  Forma incompleta Stach, 1965
Species tuberculatus Delamare Deboutteville & Massoud, 1963:201
Species tucumanensis Delamare Deboutteville & Massoud, 1963:205
Species tunicatus Salmon, JT, 1954:216
Species victorius Nguyen, 2001
Species wakhanicus Yosii, 1966
Species woringeri Delamare Deboutteville & Massoud, 1963:207

Genus Pseudokatianna Salmon, JT, 1944:179
Species aurea Salmon, JT, 1944:180
Species campbellensis Salmon, JT, 1949:52
Species fagophila Salmon, 1946
Species fasciata (Salmon, JT, 1944:178)
Species livida (Salmon, JT, 1943:2) Salmon, JT, 1944:181
Species lutea Salmon, 1946
Species minuta Salmon, 1946
Species nigra Salmon, 1946
Species nigretalba Salmon, JT, 1944:179, t.t.
Species niveovata Salmon, 1946
Species triclavata Salmon, JT, 1949:53
Species triverrucata Salmon, JT, 1944:181
Species umbrosalata Salmon, 1946
Species zebra Salmon, 1946

Genus Stenognathellus Stach, 1956
Remark by Ellis, W.N. & Bellinger, P.F. 1973:54: Stenognathellus Cassagnau, 1953 is an unavailable name (art. 13b): type species not designated.

Species cassagnaui Yosii, 1966
Species denisi Cassagnau, P, 1953:227, t.t.
Species polygonalis Hüther, 1967
Species stenognathus (Börner, 1907) Mari Mutt, JA & Bellinger, PF, 1990:144

Genus Katiannellina Delamare Deboutteville & Massoud, 1963
Species patagonica Delamare Deboutteville & Massoud, 1963:217, t.t.

Genus Betschurinus Dallai, R et Martinozzi, I, 1980:31
Species farmae Dallai, R et Martinozzi, I, 1980:33, t.t.

Genus Dalianus Cassagnau, 1969
Species galae Cassagnau, 1969

Genus Katiannina Maynard EA et Downs WL in Maynard, EA, 1951:242
Species macgillivrayi (Banks, 1897) Maynard, EA, 1951:242, t.t.
Katiannina macgillivrayi from the USA
After Maynard EA Plate XXII Fig.377a
Katiannina macgillivrayi from the USA
After Maynard EA Plate XXIX Fig.552
Katiannina macgillivrayi from the USA
After Maynard EA Plate XXIX Fig.553
Katiannina macgillivrayi from the USA
2006 © Bernard, E.C.
Katiannina macgillivrayi from the USA
2007.05.15 © Murray, T.
Katiannina macgillivrayi from the USA
2007.05.15 © Murray, T.
Katiannina macgillivrayi from the USA
2007.05.15 © Murray, T.
Katiannina macgillivrayi from the USA
2008.12.14 © Fleming, K.
Katiannina macgillivrayi from the USA
2009.05.08 © Loser, I.

Species subalpina Itoh, 2000

Genus Millsurus Betsch, 1977
Species sminthurinus (Mills, HB, 1934:99)

Genus Neokatianna Snider, RJ, 1989:541
Species acantholaema Snider, RJ, 1989:542, t.t.

Genus Rusekianna Betsch, J-M, 1977:61
Rusekianna from Norway
2009.06.03 © Fjellberg, A.
Pores at base of tibiotarsus

Species albifrons (Tullberg, 1871) Fjellberg, A, 2007:il
Sminthurinus albifrons from Norway
2006 © Fjellberg, A.
Rusekianna albifrons from France
2007.03.17 © Lebeaux, P.
Rusekianna albifrons from France
2007.03.17 © Lebeaux, P.

Species bescidica Smolis & Skarzynski, 2006
Species mongolica Betsch, J-M, 1977:62, t.t.
Species sibirica Bretfeld, 2002

Genus Zebulonia Betsch, J-M, 1970:69
Species massoudi Betsch, J-M, 1970:52

Genus Gisinianus Betsch, 1977
Species flammeolus (Gisin, 1957) Szeptycki & Weiner in Razowski, 1990:24, t.t.

Genus Vesicephalus Richards, WR in Delamare Deboutteville, C & Massoud, Z, 1964:75 key
Vesicephalus sp. from the USA
Specimen caught in spider web
2007.12.02 © Ten Eyck, C.
Remark by Delamare Deboutteville, C. & Massoud, Z. 1964:75: A French translation of the original generic diagnosis of Vesicephalus Richards, W.R. (1956:79, unpublished thesis) was published.
Remark by Bretfeld, G. 1999:98: The new monophylum Katiannida Breteld, 1986 contains the Katiannidae Börner, 1913 sensu Betsch, 1980 and the genera incertae sedis sensu Betsch, 1980 and Bretfeld, 1986, including Vesicephalus Richards in Delamare Deboutteville & Massoud, 1964, based on two diagnostic apomorphies: 1. the presence of 2 additional outer setae Ka and Kp distally at the tibiotarsus; 2. the presence of spatulate setae distally at the tibiotarsus.
Remark by Fauna Europaea 2005: Family placement is uncertain. Vesicephalus is intermediate between Katiannidae (anal appendage in females) and Sminthuridae (dorsal chaetotaxy).

Species sp.1 Richards, 1968
Species sp.2 Snider & Janssens, 2008
Vesicephalus sp. from the USA
2008.03.04 © Fleming, K.R.
Vesicephalus sp. from the USA
2008.03.04 © Fleming, K.R.

Species bellingeri Bretfeld, 2002
Species crossleyi Snider, RJ, 1985:567
Vesicephalus crossleyi from the USA
Female, type I
After Snider R, 1985 Fig.1
Vesicephalus crossleyi from the USA
Female, type I
After Snider R, 1985 Fig.2
Vesicephalus crossleyi from the USA
Female, type II
After Snider R, 1985 Fig.3
Vesicephalus crossleyi from the USA
Female, type II
After Snider R, 1985 Fig.4
Vesicephalus crossleyi from the USA
Male
After Snider R, 1985 Fig.5
Vesicephalus crossleyi from the USA
Male
After Snider R, 1985 Fig.6

Species europaeus Ardanaz & Pozo, 1985
Vesicephalus europaeus
2002 © Baquero, E. & Jordana, R.

Species longisetis (Guthrie, JE, 1903:51) Delamare Deboutteville, C & Massoud, Z, 1964:73, t.t.
Species occidentalis (Mills, HB, 1935:138) Delamare Deboutteville, C & Massoud, Z, 1964:75
Vesicephalus occidentalis from the USA
Female
After Snider R, 1985 Fig.48
Vesicephalus occidentalis from the USA
Female
After Snider R, 1985 Fig.47
Vesicephalus occidentalis from the USA
2008.11.22 © Hall, K.
Vesicephalus occidentalis from the USA
2008.11.22 © Hall, K.




References