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   Nautiloids are the only large, mobile predators which we have evidence from this period. They Orthocerasmust have enjoyed free reign of the oceans, preying most likely on the numerous trilobites. By the end of the Cambrian, this subclass of cephalopods had diversified into many different shapes and sizes. Some shells had partially uncoiled to straight shapes and ranged from a few centimeters to 3 meters across. There are even fragments that suggest sizes up to 10m! For 100 million years the nautiloids must have been the rulers of the oceans, until the development of a similar but more successful creature: ammonoids.
      At first glance, it is amazing that a different classification was given to these other shelled cephalopods. There are however, several subtle, but significant differences. Two major contrasting features stand out once the inside of the phragmacone is observed. The siphuncle, which in nautiloids is well within the interior of the shell, is instead placed next to the outer surface. More significantly, the sutures (the area where the septa meet the inner wall of the shell) are not straight as in the Nautilus. They instead trace complex, waving patterns. This is the main characteristic that divides the two classifications, though the actual function of complex sutures is still far from resolved and an area of debate among invertebrate paleontologists.
 

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