http://www.collembola.org/taxa/onycinae.htm - Last updated on 2009.09.14 by Frans Janssens
Checklist of the Collembola: Onychiurinae

Subfamilia Onychiurinae Börner, 1901 key
Genus Number
of species
Absolonia1
Agraphorura12
Allonychiurus22
Archaphorura3
Argonychiurus6
Arneria2
Bionychiurus3
Chribellphorura1
Cribrochiurus2
Deharvengiurus9
Detriturus4
Deuteraphorura71
Dimorphaphorura1
Dinochiurus1
Heteraphorura15
Hymenaphorura41
Kalaphorura10
Megaphorura2
Micraphorura15
Micronychiurus6
Oligaphorura18
Ongulonychiurus1
Onychiuroides25
Onychiurus47
Orthonychiurus29
Paronychiurus7
Pilonychiurus1
Probolaphorura2
Protaphorura127
Protaphorurodes1
Psyllaphorura8
Reducturus1
Sacaphorura1
Sensillonychiurus3
Similonychiurus1
Spelaphorura1
Spinonychiurus3
Supraphorura1
Tantulonychiurus2
Thalassaphorura36
Uralaphorura5
Vexaphorura1
Vibronychiurus5
Wandaphorura1
Yoshiiphorura1
Onychiurinae555
Genera with number of species
Onychiurinae from China
2008.07.17 © NCode, A.
Onychiurinae from Canada
2009.01.09 © Weeta, W.


Tribus Hymenaphorurini Pomorski RJ, 1996:99

Genus Kalaphorura Absolon, K, 1901:388habitus dorsal
Species bearei (Bagnall, 1937) Salmon, JT, 1959:151
Species bermani (Martynova, 1976) Weiner, WM, 1996:179
Species burmeisteri (Lubbock, J, 1873:190) Handschin, 1920, t.t., B
Lipura Burmeisteri(sic) from the UK
After Lubbock, J., 1873 Pl.44.
Remark by Dányi, L. & Traser, G. 2008:22: According to Dunger (1975) the early Hungarian data of Kalaphorura burmeisteri could include Kalaphorura paradoxus (Schäffer, 1900) as well.

Kalaphorura burmeisteri from the UK
Vestigial furca
2002 © Hopkin, S.P.
Kalaphorura burmeisteri from the UK
2004 © Hopkin, S.P.
Kalaphorura burmeisteri from the UK
2004 © Hopkin, S.P.
Kalaphorura burmeisteri from the UK
2006 © Valentine., B
Kalaphorura burmeisteri from the UK
2006 © Barton, T.
Kalaphorura burmeisteri from the UK
2006 © Cornwall, N.J.
Kalaphorura burmeisteri from Belgium
2007 © Doornaert, A.
Kalaphorura burmeisteri from the UK
2007 © Tonsbeek, M.
Kalaphorura burmeisteri from the UK
2007 © Tonsbeek, M.
Kalaphorura burmeisteri from the UK
2007.11.28 © Oakshott, A.
Kalaphorura burmeisteri from the UK
2007.11.28 © Oakshott, A.
Kalaphorura burmeisteri from the UK
2008.12.24 © Barton, T.
Kalaphorura burmeisteri from the UK
Large anals spines
2008.12.24 © Barton, T.
Kalaphorura burmeisteri from the UK
Tuberculate cuticula
2009.01.01 © Barton, T.

Species carpenteri (Stach, 1919) Handschin, 1924
Species granulata (Börner, C, 1909:103) Handschin, 1924
Species heterodoxa (Gisin, H, 1964:655) Weiner, WM, 1996:179
Species ludzaki Weiner, WM et Stomp, N, 2005:96
Species paradoxa (Schäffer, 1900) Ellis, WN & Bellinger, PF, 1973:26, B
Remark by Dányi, L. & Traser, G. 2008:22: Loksa's "Onychiurus burmeisteri" data were corrected by Traser (1999) as Kalaphorura paradoxa.

Species tuberculata (Moniez, 1890) Gruia, 1964
  Subspecies clavipila Stach, 1920
Species vannieri (Massoud, 1968) Weiner, WM, 1996:179

Genus Hymenaphorura Bagnall, 1948 key
Species alaskana Pomorski, RJ, 2001:451
Species alpina (Stach, J, 1946:8) Salmon, JT, 1964:159
Species alticola (Bagnall, 1935) Salmon, JT, 1959:153
Species anatolii Pomorski, RJ, 2001:467
Species arantiana Weiner & Stomp, 2001
Hymenaphorura arantiana from Luxembourg
After Stomp N & Weiner WM, 2005 Fig.6

Species californica (Coleman, 1941, i.s.) Salmon, 1964:159
Species cocklei (Folsom, 1908) Salmon, JT, 1964:159
Hymenaphorura cocklei from the USA
2008.07.07 © Wagner., B
Hymenaphorura cocklei from the USA
2008.12.28 © Hampel, P.
Hymenaphorura cocklei from the USA
2008.12.28 © Hampel, P.
Hymenaphorura cocklei from the USA
2009.01.13 © Davis, J.
Hymenaphorura cocklei from the USA
2009.01.13 © Davis, J.

Species deca (Christiansen, K & Bellinger, P, 1980:431) Pomorski, RJ, 2001:447
Species dentifera (Stach, 1934) Salmon, 1964:161
Remark by Dányi, L. & Traser, G. 2008:21: Hymenaphorura dentifera (Stach, 1934) is a "species inquirenda according to Gisin (1960)". [Janssens, F., 2008.11.17: Contrary to what Dányi & Traser (2008:21) claim, Gisin (1960:151) refers to fimetarius var. dentifera Kos, 1939, not to sibiricus f. dentifera Stach, 1934. And the first is considered as a species dubia by Gisin (1960:151), not as species inquirenda.]

Species gamae Arbea, JI et Jordana, R in Jordana, R et Ariño, AH, 1994:50
Species granulata Pomorski, RJ, 2001:453
Species hispanica Pomorski, 1992
Species improvisa Pomorski & Skarzynski, 2000
Species jugoslavica (Gisin, 1963) Pomorski, RJ, 2002:il
Species liberta Pomorski, RJ, 1990:219
Species magnina Wray, 1950, i.s.
Species maiteae Arbea, JI et Jordana, R in Jordana, R et Ariño, AH, 1994:51
Species montana (Handschin, 1921) Salmon, JT, 1964:159
Species mystica Pomorski, RJ, 2001:462
Species nearctica Pomorski, RJ, 2001:460
Species nicolae Barra, J-A, 1998:522
Species nova Pomorski, RJ, 1990:210
Species palaearctica Pomorski, RJ, 2001:467
Species parva Skarzynski, D et Pomorski, RJ, 1996:319
Species polonica Pomorski, RJ, 1990:213
Species pseudosibirica (Stach, 1954:101) Salmon, JT, 1964:160
Species rafalskii Weiner, WM et Szeptycki, A, 1997:193
Species reducta Pomorski, RJ, 2001:442
Species reisingeri (Neuherz, 1979) Pomorski, RJ, 2002:il
Species ridibunda Pomorski, RJ, 2001:451
Species sensitiva Pomorski, RJ, 2001:457
Species sibirica (Tullberg, T, 1877:40) Bagnall, 1948:637, t.t., B
Species similis (Folsom, 1917) Bagnall, 1948
Species strasseri (Stach, 1934) Salmon, 1964:161
Species submontana (Denis, 1926) Salmon, 1964:161
Species subsimilis (Bagnall, 1948) Salmon, JT, 1959:145
Species superba Pomorski, RJ, 2001:457
Species teretis Pomorski, RJ, 2001:464
Species troglodytes Bagnall, 1948
Species uzicensis Curcic, BPM, Lucic, LR, Curcic, SB et Curcic, NB, 2005:176
Species valdegranulata (Stach, 1954:103) Salmon, 1964:161

Genus Heteraphorura Bagnall, 1948 key
Species bima (Christiansen, K & Bellinger, P, 1980:438) Pomorski, RJ, 2002:461
Species carpatica (Stach in Kseneman, 1938:468) Weiner, WM, 1996:177
Species casa (Christiansen, K & Bellinger, P, 1980:440) Pomorski, RJ, 2002:463
Species conjungens (Börner, C, 1909:104) Pomorski, RJ, 2002:459
Species edentata (Kos, 1939) Pomorski, RJ, 2002:459
Species intricata Pomorski, RJ, 2002:468
Species japonica (Yosii, 1967) Pomorski, RJ, 2002:459
  Subspecies kyotensis Yoshii, 1996
Species justynae Pomorski, RJ, 2002:465
Species longisetosus (Lee & Park, 1986) Pomorski, RJ, 2002:459
Species orientalis (Martynova, 1976) Pomorski, RJ, 2002:459
Species pseudoseolagensis (Martynova, 1981) Pomorski, RJ, 2002:459
Species seolagensis (Lee, 1974) Pomorski, RJ, 2002:459
Species subtenuis (Folsom, 1917) Pomorski, RJ, 2002:459
Species tala (Christiansen, K & Bellinger, P, 1980:444) Pomorski, RJ, 2002:461
Species variotuberculata (Stach, J, 1934:161) Bagnall, 1948 tt

Genus Arneria Pomorski, 2000
Species alinae Pomorski, 2000
Species filiformis Pomorski, 2000

Genus Paronychiurus Bagnall, 1948 key
Species dentatus (Folsom, JW, 1902:91), i.s.
Species eous (Christiansen, K & Bellinger, P, 1980:449) Pomorski, RJ, 2003:190
Species hubbardi Pomorski, RJ, 2003:192
Species imadatei (Yosii, R, 1956:58, i.s.) Salmon, JT, 1964:187
Species japonicus (Yosii, R, 1956:57) Salmon, JT, 1964:186
Species probus Pomorski, RJ, 2003:191
Species ramosus (Folsom, 1917) Bagnall, 1948, t.t.

Genus Psyllaphorura Bagnall, 1948
Species bashkirica (Khanislamova, 1986) Weiner, WM, 1996:181
Species jiangsuensis Yan, HJ, Huang, C et Chen, J-X, 2007:?
Species martynovae (Stebaeva, 1985, s.d.) Weiner, WM, 1996:181
Species obesa (Mills, HB, 1934:35) Bagnall, 1948, t.t.
Species okafujii (Yosii, 1967) Weiner, WM, 1996:181
Species ryozoyoshii Weiner & Najt, 2000
Species sensillifera (Martynova, 1981) Weiner, WM, 1996:181
Species uenoi (Yosii, R, 1954:62) Weiner, WM, 1996:181

Genus Dinochiurus Pomorski, RJ et Steinmann, D, 2004:15
Species reus (Christiansen, K & Bellinger, P, 1980:433) Pomorski, RJ & Steinmann, D, 2004:17, t.t.
Dinochiurus reus from the USA
After Pomorski RJ & Steinmann D, 2004 Fig.1


Genus Reducturus Pomorski, RJ et Steinmann, D, 2004:19
Species christianseni Pomorski, RJ et Steinmann, D, 2004:19, t.t.
Reducturus christianseni from the USA
After Pomorski RJ & Steinmann D, 2004 Fig.15


Genus Sacaphorura Pomorski, RJ et Steinmann, D, 2004:20
Species parapseudocellata Pomorski, RJ et Steinmann, D, 2004:23, t.t.
Sacaphorura parapseudocellata from the USA
After Pomorski RJ & Steinmann D, 2004 Fig.19-20


Genus Vexaphorura Pomorski, RJ et Steinmann, D, 2004:23
Species robustiseta Pomorski, RJ et Steinmann, D, 2004:23, t.t.
Vexaphorura robustiseta from the USA
After Pomorski RJ & Steinmann D, 2004 Fig.29


Genus Wandaphorura Pomorski, RJ, 2007:53
Species millsi (Chamberlain, 1943) Pomorski, RJ, 2007:53, t.t.

Genus Probolaphorura Dunger, W, 1977:1
Remark by Pomorski, R.J. & Steinmann, D. 2004:26: The tribe Hymenaphorurini probably also comprises the genus Probolaphorura, but it needs redescription.

Species plumosetosa Martynova, 1978
Species sachalinensis Dunger, W, 1977:2, t.t.

Genus Protaphorurodes Bagnall, 1949
Species tomuraushensis (Yosii, 1940) Pomorski, RJ, 2002:il

Tribus Oligaphorurini Bagnall, 1949

Genus Archaphorura Bagnall, 1949
Species alavensis (Simón-Benito, JC & Luciáñez, MJ in Jordana, R & Ariño, AH, 1994:73) Pomorski, RJ, 2002:il
Species marcuzzii Cassagnau, 1968
Species serratotuberculata (Stach, 1933) Bagnall, 1949, t.t.

Genus Oligaphorura Bagnall, 1949
Species aborigensis (Fjellberg, 1987) Pomorski, RJ, 2002:il
Species gela (Christiansen, K & Bellinger, P, 1980:407) Pomorski, RJ, 2002:il
Species groenlandica (Tullberg, T, 1877:41) Pomorski, 1996:31, t.t.
Onychiurus groenlandicus from Greenland
After Folsom JW, 1919 Fig.14

Species judithae (Weiner, 1994) Pomorski, RJ, 1996:91
Species koreana (Weiner, 1994) Weiner, WM, 1996:176
Species kurtshevae Martynova, 1981
Species linderae (Weiner, 1994) Weiner, WM, 1996:176
Species melittae Christian, E, 1993:163
Species montana Weiner, 1994
Species nuda (Fjellberg, 1987) Pomorski, RJ, 2002:il
Species palissai (Yosii, 1971) Pomorski, RJ, 2002:il
Species pingicola (Fjellberg, 1987) Pomorski, RJ, 2002:il
Species reversa (Fjellberg, 1987) Pomorski, RJ, 2002:il
Species sabulosa Babenko, AB, 2007:366
Species schoetti (Lie-Pettersen, 1896) Bagnall, 1949
Oligaphorura schoetti from the UK
2002 © Hopkin, S.P.

Species tottabetsuensis (Yosii, R, 1972:83) Pomorski, RJ, 2002:il
Species tuvinica Potapov & Stebaeva, 1997
Species ursi Fjellberg, 1984

Genus Micraphorura Bagnall, 1949 key
Species absoloni (Börner, 1901) Pomorski, 1996:35, t.t., B
Species alna (Fjellberg, 1987) Weiner, WM, 1996:175
Species chatyrdagi Kaprus, IJ, Weiner, WM et Pomorski, RJ, 2002:356
Micraphorura chatyrdagi from Ukraine
After Kaprus I.J., Weiner, W.M. & Pomorski, R.J, 2002 Fig.16

Species daii Pomorski, RJ, Skarzynski, D et Kaprus, I, 1998:253
Species eremia Kaprus, IJ, Weiner, WM et Pomorski, RJ, 2002:354
Micraphorura eremia from Ukraine
After Kaprus I.J., Weiner, W.M. & Pomorski, R.J, 2002 Fig.9

Species hackeri (Christian, E, 1986:177) Christian, E, 2002:264
Species interrupta (Fjellberg, 1987) Pomorski, RJ, 2002:il
Species irinae Thibaud & Taraschuk, 1997
Species multiperforata (Gruia, M, 1973:173) Pomorski, RJ, 2002:il
Species nataliae (Fjellberg, 1987) Kaprus, IJ, Weiner, WM & Pomorski, RJ, 2002:356
Species pieninensis Weiner, 1988
Species pseudoraxensis (Nosek, J & Christian, E, 1983:397) Christian, E, 2002:264
Species raxensis (Gisin, 1961) Christian, E, 2002:264
Species steposa Kaprus, IJ, Weiner, WM et Pomorski, RJ, 2002:353
Micraphorura steposa from Ukraine
After Kaprus I.J., Weiner, W.M. & Pomorski, R.J, 2002 Fig.1

Species uralica (Khanislamova, 1986) Kaprus, IJ, Weiner, WM & Pomorski, RJ, 2002:353

Genus Chribellphorura Weiner, WM, 1996:174
Species allanae (Christiansen, K & Bellinger, P, 1980:405) Weiner, WM, 1996:174, t.t.

Genus Dimorphaphorura Bagnall, 1949
Species quadrituberculata (Börner, 1901) Pomorski, RJ, 2002:il

Tribus Protaphorurini Bagnall, 1949

Genus Supraphorura Stach, 1954
Species furcifera (Börner, 1901) Pomorski, 1996:27, B
Supraphorura furcifera from the UK
Vestigial furca
2002 © Hopkin, S.P.


Genus Protaphorura Absolon, K, 1901:387 key
Protaphorura sp. from Germany
2006.12.30 © Kohl, F.

Species aconae Arbea, JI et Jordana, R in Jordana, R et Ariño, AH, 1994:43
Species ajudagi Pomorski, RJ, Skarzynski, D et Kaprus, I, 1998:259
Species aksuensis (Martynova, 1972) Pomorski, RJ, 2002:il
Species armata (Tullberg, 1869) Börner, 1909, t.t., B
Onychiurus armatus from New Zealand
After Salmon, J.T., 1941 Pl.47 Fig.125.
Protaphorura armata from the UK
Vestigial furca
2002 © Hopkin, S.P.
Protaphorura armata from Sweden
2005 © Hall, K.
 
Protaphorura armata from Sweden
2005 © Hall, K.
Protaphorura armata from Denmark
2007.02.09 © Larsen, T.
Protaphorura armata from the UK
2007.03.18 © Cornwall, N.J.
Protaphorura armata from France
2008.mm.dd © Lebeaux, P.
Protaphorura armata from the UK
2008.04.07 © Farrell, S.
Protaphorura armata from the UK
2008.04.08 © Farrell, S.
Protaphorura armata from Tasmania
2009.08.16 © Daley, A.

  Subspecies denticulata Handschin, 1924
  Subspecies furcifera Schött, 1924
  Subspecies inermis Axelson, WM, 1905:790
  Subspecies islandica Gisin, 1960
  Subspecies trispina Axelson, 1935
  Forma occidentalis Pomorski RJ, 1998:95
  Forma scanica Bödvarsson, H, 1970:6
  Forma silesiaca Pomorski RJ, 1998:95
Species asensitiva (Stach, 1954) Salmon, JT, 1964:170
Species aurantiaca (Ridley, 1880) Salmon, JT, 1964:164
Onychiurus aurantiacus from the UK
After Pitkin, B.R., 1980 Fig.1
Protaphorura aurantiaca from the UK
2008.02.26 © Botting, J.
Protaphorura aurantiaca from the UK
2008.04.11 © Robertson, A.
Protaphorura aurantiaca from the UK
2009.01.24 © Robertson, A.
Protaphorura aurantiaca from the UK
2009.04.12© Robertson, A.

Species austriaca (Butschek, 1948) Salmon, JT, 1964:170
Species bicampata (Gisin, 1956) Salmon, JT, 1964:170
Species boedvarssoni Pomorski, RJ, 1993:125
Species borealis Martynova, 1973
Species borzica (Gruia, 1999) Pomorski, Leithner & Bruckner, 2003
Species boskovae Lucic, Dimitrijevic & Mihajlova, 2003
Species brevispinata (Yosii, R, 1966:548) Pomorski, RJ, 2002:il
Species caerulea (Börner, 1901) Dallai & Malatesta in Calderini, 1995:7
Species caledonica (Bagnall, 1935) Kloet, GS & Hincks, WD, 1945:5
Species campata (Gisin, 1952) Salmon, JT, 1964:170, B
Species cancellata (Gisin, 1956) Salmon, JT, 1964:170
Species chilensis (Loksa & Rubio, 1966) Pomorski, RJ, 2002:il
Species christiani Pomorski, RJ, Leithner, C et Bruckner, A, 2003:307
Species churchilliana (Hammer, 1953) Salmon, JT, 1964:170
Species crassa (Kos, 1940) Salmon, 1964:169
Species dallaii Nosek & Paoletti, 1981
Species daviesi (Bagnall, 1935) Kloet, GS & Hincks, WD, 1945:5
Species decempunctata (Kos, 1939) Salmon, JT, 1964:170
Species diplosensillata (Dunger, 1978)
Species discrepans (Bagnall, 1948) Salmon, JT, 1959:154
Species dudichi (Loksa, I & Rubio, I, 1966:148) Dallai, R, Malatesta, E & Ramellini, P in Minelli, A, Ruffo, S & La Posta, S, 1995:7
Species duodecimpunctata (Folsom, JW, 1919:6A) Salmon, JT, 1964:171
Remark by Folsom, J.W. 1919:6A: Pseudocelli of antennal bases 6+6 (two specimens) or 5+5 (one specimen).
Onychiurus duodecimpunctatus from Arctica
After Folsom JW, 1919 Plate 3 Fig.29
Remark by Hammer, M. 1953:36-37: In specimens from Northern Canada, the number of pseudocelli at the antennal base of Onychiurus 12-punctatus(sic) varies between 4 to 6. Of 23 specimens, 10 were found with 4+4, 11 with 5+5, 1 with 6+6 and 1 with a bilateral asymmetric number of 5+4 pseudocelli.

Remark by Janssens, F. 2008.08.15: Given the description of 'decimpunctatus Skidmore, 1995:53' is lacking it is a nomen nudum, therefore an invalid name. The name implicitly refers to specimens of duodecimpunctatus having 5 pseudocelli at the antennal base in steadt of the 6 pseudocelli of the nominal form. Such specimens are considered to be an intraspecific variation of duodecimpunctatus. Therefore, 'decimpunctatus Skidmore, 1995:53' is a partial junior synonym of duodecimpunctatus Folsom, 1919:6A.


Species edentata (Kos, 1939) Salmon, JT, 1964:174
Species eichhorni (Gisin, 1954) Salmon, JT, 1964:171, B
Protaphorura eichhorni from Luxembourg
After Stomp N & Weiner WM, 2005 Fig.11

Species elenae Kaprus, IJ et Pomorski, RJ, 2008:668
Protaphorura elenae from Russia
After Kaprus IJ & Pomorski RJ, 2008 Fig.1

Species fimata (Gisin, 1952) Salmon, JT, 1964:171
Species fistania Thibaud & Peja, 1994
Species florae Simón-Benito, JC et Luciáñez, MJ in Jordana, R et Ariño, AH, 1994:70
Species fredericoi Arbea, JI, 2003:93
Species gisini (Haybach, 1960) Selga & al., 1977
Species glebata (Gisin, 1952) Dallai & Malatesta in Calderini, 1995:8
Species ianstachi Yosii, 1972
Species ionescui Radwanski & al., 2006
Species jacutica Martynova, 1976
Species janosik Weiner, 1990
Species januarii (Weiner, 1977) Pomorski, RJ, Leithner, C & Bruckner, A, 2003:309
Species kadici (Loksa, I, 1967:290) Pomorski, RJ, 2002:il
  Varietas geminiocellatus Loksa, I, 1967:292
Species karadzicae Lucic, Dimitrijevic & Mihajlova, 2003
Species kaszabi Dunger, 1978
Species kolenatii Hemmer, 2000
  Forma alba Hemmer, 2000
  Forma rubra Hemmer, 2000
Species kopetdagi Pomorski, RJ, 1994:193
Species kultia (Baijal, 1956) Salmon, JT, 1964:172
Species latosimilis (Loksa, I et Rubio, I, 1966:149)
Species levantina (Christiansen, K, 1956:123) Salmon, JT, 1964:172
Species licheniphila Kaprus, IJ et Pomorski, RJ, 2008:670
Protaphorura licheniphila from Russia
After Kaprus IJ & Pomorski RJ, 2008 Fig.6

Species longisensillata (Yosii, 1969)
Species macfadyeni (Gisin, 1953) Salmon, JT, 1964:172
Protaphorura macfadyeni from Russia
After Kaprus IJ & Pomorski RJ, 2008 Fig.11

Species macrodentata (Hammer, 1953) Salmon, JT, 1964:172
Species macrotuberculata (Kos, 1939, i.s.) Salmon, JT, 1964:172
Species maglicensis Zivadinovic, 1970
Species matsumotoi (Kinoshita, 1923) Pomorski, RJ, 2002:il
Species mediovanderdrifti (Haybach, 1960) Pomorski, RJ, 2002:il
Species meridiata (Gisin, 1952) Salmon, JT, 1964:172
Species merita Kaprus, IJ et Pomorski, RJ, 2008:672
Protaphorura merita from Russia
After Kaprus IJ & Pomorski RJ, 2008 Fig.16-17

Species metapulvinata (Loksa, I, 1964:91) Pomorski, RJ, 2002:il
Species microcellata Dunger, 1978
Species microtica Dunger, 1978
Species mongolica (Martynova, 1975) Pomorski, RJ & Kaprus, I, 2007:137
Protaphorura mongolica from Mongalia
After Kaprus IJ & Pomorski RJ, 2008 Fig.24

Species montana Mateos Frias, EM et Arbea, JI, 1986:283
Species multipunctata (Absolon, K, 1900:406, i.s.) Salmon, JT, 1964:169
Species multituberculata (Stach, J, 1934:127) Pomorski, RJ, 2002:il
Species nazarovensis Kaprus, IJ et Pomorski, RJ, 2008:678
Protaphorura nazarovensis from Russia
After Kaprus IJ & Pomorski RJ, 2008 Fig.28

Species neriensis Martynova, 1976
Species nutak (Yosii, R, 1972:86) Pomorski, RJ, 2002:il
Protaphorura nutak from Russia
After Kaprus IJ & Pomorski RJ, 2008 Fig.32

Species octopunctata (Tullberg, T, 1877:40) Kloet, GS & Hincks, WD, 1945:5, B
Protaphorura octopunctata from Russia
After Pomorski RJ & Kaprus IJ, 2007 Fig.1

Species okaensis (Becker, 1905, i.s.) Salmon, JT, 1964:173
Species ombrophila (Stach, J, 1960:509) Pomorski, RJ, 2002:il
Species orthacantha (Handschin, 1920) Salmon, JT, 1964:173, B
Species pannonica (Haybach, 1960) Pomorski, 1990
Species parallata (Gisin, 1952) Salmon, JT, 1964:174
Species paranemorata (Selga, 1962) Pomorski, RJ, 2002:il
Species paucisetosa (Hammer, 1942) Skidmore, 1995:53
Species pjasinae (Martynova, 1976) Pomorski, RJ, 2002:il
Species pseudarmata (Folsom, 1917) Salmon, JT, 1964:174
Species pseudocellata (Naglitsch, 1962)
Species pseudoglebata Arbea, JI et Jordana, R, 1989:11
Species pseudostyriaca (Loksa, I, 1964:88) Pomorski, RJ, Leithner, C & Bruckner, A, 2003:307
Species pseudovanderdrifti (Gisin, 1957) Salmon, JT, 1964:174
Species quadriocellata (Gisin, 1947) Salmon, JT, 1964:170, B
Protaphorura quadriocellata from Europe
After Pomorski RJ & Kaprus I, 2007 Fig.11

Species quercetana Mateos Frias, EM et Arbea, JI, 1986:285
Species saltuaria Pomorski, RJ et Kaprus, I, 2007:135
Protaphorura saltuaria from Ukraine
After Pomorski RJ & Kaprus I, 2007 Fig.16

Species sakatoi (Yosii, 1966)
Species sensillata (Khanislamova, 1986) Pomorski, RJ, 2002:il
Species septempapillata Palissa, 1986
Species sp. Kaprus, IJ, Pomorski, RJ, Skarzynski, D et Potapov, MB, 2005:893
Species spinoidea (Steiner, 1955) Salmon, JT, 1964:175
Species stachi (Bagnall, 1935) Kloet, GS & Hincks, WD, 1945:5
Remark by Pitkin.R., B 1980:423: Type specimen not adult.

Species stalagmitorum (Absolon, K, 1900:407) Pomorski, RJ, 2002:il
Species stiriaca (Stach, J, 1946:13) Salmon, JT, 1964:175
Species stogovi Pomorski, RJ, 1993:121
Species strasseri Stach, J, 1934:147
Species subaequalis (Bagnall, 1937) Kloet, GS & Hincks, WD, 1945:5
Remark by Pitkin.R., B 1980:423: Type specimen not adult.

Species subarctica Martynova, 1976
Species subcancellata (Gisin, 1963) Pomorski, RJ, 2002:il
Species subfimata Thibaud & Christian, 1986
Species submersa Kaprus, IJ et Pomorski, RJ, 2008:681
Protaphorura submersa from Russia
After Kaprus IJ & Pomorski RJ, 2008 Fig.37

Species subnemorata (Gisin, 1957) Salmon, JT, 1964:175
Species suboctopunctata Khanislamova, 1986
Species subparallata (Selga, 1962) Pomorski, RJ, 2002:il
Species svontoernei (Gisin, 1957) Salmon, JT, 1964:175
Remark by Nicolson, D. 2007.mm.dd:in litt.: Based on art.NN the hyphen in the original spelling is removed.

Species taimyrica Martynova, 1976
Species tamdaona Nguyen, 1995
Species teres (Yosii, R, 1956:53) Salmon, JT, 1964:174
Species tetragrammata (Gisin, 1964) Kaprus, IJ & Pomorski, RJ, 2008:684
Protaphorura tetragrammata from Russia
After Kaprus IJ & Pomorski RJ, 2008 Fig.47

Species tolae Pomorski, RJ et Kaprus, I, 2007:131
Protaphorura tolae from Russia
After Pomorski RJ & Kaprus I, 2007 Fig.6

Species tricampata (Gisin, 1956) Salmon, JT, 1964:175
Species tridentata Stebaeva, 1982
Species triparallata (Gisin, 1960) Salmon, JT, 1964:175
Species trivontoernei (Gisin, 1957) Salmon, JT, 1964:175
Species troglophila Nosek, 1975
Species tschernovi Martynova, 1976
Species tullbergi (Bagnall, 1937) Kloet, GS & Hincks, WD, 1945:5
Species tundricola Martynova, 1976
Species unari Rusek, 1995
Species ussurica Martynova, 1981
Species valsainensis Acón, 1981
Species vontoernei (Gisin, 1957) Salmon, JT, 1964:176
Species yagii (Miyoshi, 1923) Salmon, JT, 1964:174
Species zlatiborensis Lucic, Curcic, Pavkovic-Lucic & Tomic, 2008
Species zloti Curcic & Lucic, 1997
Species zori Martynova, 1975

Genus Spelaphorura Bagnall, 1948
Species glenniei Bagnall, 1948, t.t.

Genus Yoshiiphorura Jordana R et Martínez M in Martínez, M, Baquero, E, Barranco, P, Ariño, AH et Jordana, R, 2004:3
Species bellingeri Jordana R et Martínez M in Martínez, M, Baquero, E, Barranco, P, Ariño, AH et Jordana, R, 2004:7, t.t.

Genus Megaphorura Fjellberg, A, 1998:108
Species arctica (Tullberg, T, 1877:39) Fjellberg, A, 1998:108, t.t.
Megaphorura arctica from the UK
2002 © Hopkin, S.P.
Megaphorura arctica from the Arctic region
2003.03.06 © Holmstrups, M.
Megaphorura arctica from the Arctic region, dehydrated
2003.03.06 © Holmstrups, M.
Megaphorura arctica from the Arctic region, hydrated
2007 © Borland, R.
Megaphorura arctica from the Arctic region, dehydrated
2007 © Borland, R.
Megaphorura arctica from Sweden
2007.12.03 © Hall, K.

Species evansi Bagnall, 1935
Remark by Pitkin.R., B 1980:423: This species lacks a rudimentary furca and belongs to the arcticus group.


Tribus Thalassaphorurini Pomorski RJ, 1998:134

Genus Thalassaphorura Bagnall, 1949
Species alborufescens (Vogler, 1895) Pomorski, RJ, 2002:il
Species bagnalli (Salmon, JT, 1959:155) Pomorski, RJ, 2002:il
Species butrosi (Christiansen, K, 1956:126) Pomorski, RJ, 2002:il
Species clayae (Salmon, 1958) Pomorski, RJ, 2002:il
Species cryptopyga (Denis, 1931) Pomorski, RJ, 2002:il
Species debilis (Moniez, 1889) Fjellberg, A, 1998:109
Thalassaphorura debilis from the UK
2002 © Hopkin, S.P.

Species dinghuensis (Lin & Xia, 1985) Pomorski, RJ, 2002:il
Species duplopunctata (Strenzke, 1954) Pomorski, RJ, 2002:il
Species encarpata (Denis, 1931) Fjellberg, A, 1998:111
Species franzi (Stach, J, 1946:11) Pomorski, RJ, 2002:il, t.t.
Species ghatensis (Prabhoo, 1971) Pomorski, RJ, 2002:il
Species guineaensis (Salmon, 1965) Pomorski, RJ, 2002:il
Species halophila (Bagnall, 1937) Pomorski, RJ, 2002:il
Thalassaphorura halophila from the UK
2002 © Hopkin, S.P.

Species hera (Christiansen, K & Bellinger, P, 1980:419) Pomorski, RJ, 2002:il
Species hoguei (Palacios-Vargas, JG & Díaz, M, 1996:10) Pomorski, RJ, 2002:il
Species ishikawai (Yosii, 1956) Pomorski, RJ, 2002:il
Species jailolonis (Yoshii, R & Suhardjono, YR, 1992:9) Pomorski, RJ, 2002:il
Species kirgisica (Martynova, 1971) Pomorski, RJ, 2002:il
Species kwona (Thibaud & Lee, 1994) Pomorski, RJ, 2002:il
Species lagunensis (Palacios-Vargas, JG & Díaz, M, 1996:2) Pomorski, RJ, 2002:il
Species lifouensis (Thibaud, J-M & Weiner, WM in Najt, J & Matile, L, 1997:72) Pomorski, RJ, 2002:il
Species martynovae Dunger, 1978
Species moniezi (Bagnall, 1935) Pomorski, RJ, 2002:il
Remark by Pitkin.R., B 1980:423: Type specimen not adult.

Species orientalis (Stach, J, 1964:9) Pomorski, RJ, 2002:il
Species parvicornis (Mills, HB, 1934:33) Pomorski, RJ, 2002:il
Species petallata (Salmon, JT, 1958:709) Pomorski, RJ, 2002:il
Species petaloides (Rusek, 1981) Pomorski, RJ, 2002:il
Species qixiaensis Yan, Shi & Chen, 2006
Species saccardyi (Denis, JR, 1935:230) Pomorski, RJ, 2002:il
Species tamurai (Yue, Q-Y & Yin, W-Y, 2000:44) Pomorski, RJ, 2002:il
Species tenuis Babenko, AB, 2007:369
Species thaibinhensis Nguyen, 2001
Species tovtrensis (Kaprus, IJ & Weiner, WM, [1995]:61) Pomorski, RJ, 2002:il
Allaphorura tovtrensis from Ukraine
After Kaprus I.J. & Weiner W.M., [1995] Fig.9

Species yodai (Yosii, 1966) Pomorski, RJ, 2002:il
Species yolandae (Izarra, 1971) Pomorski, RJ, 2002:il
Species zschokkei (Handschin, 1919) Pomorski, RJ, 1998:139
  Varietas armata Handschin, 1924

Genus Detriturus Pomorski, RJ, 1998:142
Species jubilarius (Gisin, 1957) Pomorski, RJ, 1998:143, t.t.
Species lotius (Palacios-Vargas, JG & Díaz, M, 1996:5) Pomorski, RJ, 2002:il
Species cassagnaui (Gers, 1980) Pomorski, RJ, 2002:il
Species trilobatus (Palacios-Vargas, JG & Díaz, M, 1996:7) Pomorski, RJ, 2002:il

Genus Sensillonychiurus Pomorski, RJ et Sveenkova, YB, 2006:191 key
Species geminus Pomorski, RJ et Sveenkova, YB, 2006:195
Species minusculus Pomorski, RJ et Sveenkova, YB, 2006:193, t.t.
Species virginis Pomorski, RJ et Sveenkova, YB, 2006:193

Genus Spinonychiurus Weiner, WM, 1996:188
Spinonychiurus epaphius from Ukraine
After Kaprus' IJ & Tsalan JV, 2009 Fig.1.6
Abdominal sternum III divided into 2 subsegments

Species edinensis (Bagnall, 1935) Weiner, WM, 1996:188, t.t.
Allonychiurus edinensis from the UK
Anal spines; 4 stout setae on abd.5
2002 © Hopkin, S.P.

Species epaphius Kaprus, IJ et Tsalan, JV, 2009:174
Spinonychiurus epaphius from Ukraine
After Kaprus' IJ & Tsalan JV, 2009 Fig.1.1

Species subedinensis (Arbea, JI & Jordana, R, 1985:66) Kaprus, IJ & Tsalan, JV, 2009:176

Genus Tantulonychiurus Pomorski, 1996
Species asiaticus Babenko, AB, 2007:366
Species volinensis (Szeptycki, 1964) Pomorski, RJ, 1996:94, t.t.

Genus Agraphorura Pomorski, RJ, 1998:150 key
Species acuitlapanensis (Palacios-Vargas, JG & Deharveng, L, 1982:3) Pomorski, RJ, 2004:76
Species calvoi Arbea, JI, 2005:21
Species eisi (Rusek, 1976) Pomorski, RJ, 2004:76
Species fernandae (Oliveira & Thibaud, 1992) Pomorski, RJ, 2004:76
Species gambiaria (Murphy, 1965) Pomorski, RJ, 2004:76
Species mariapetrae (Thibaud, 1993) Pomorski, RJ, 2004:76
Species naglitschi (Gisin, 1960) Pomorski, RJ, 1998:150, t.t.
Species portucalensis (da Gama, 1964) Pomorski, RJ, 2004:76
Species pseudojusti (Thibaud & Massoud, 1980) Pomorski, RJ, 2004:76
Species sangelensis Kaprus, IJ et Stebaeva, SK, 2006:351
Agraphorura sangelensis from Southern Siberia
After Kaprus I.J. & Stebaeva S.K, 2006 Fig.1

Species spelaea Pomorski, RJ, 2004:76
Species xenonis (Ellis, WN, 1976:234) Pomorski, RJ, 2004:76

Genus Micronychiurus Bagnall, 1949
Species aranzadii (Beruete, E, Arbea, JI & Jordana, R in Jordana, R & Ariño, AH, 1994:29) Weiner, WM, 1996:186
Species bernardoi (Beruete, E, Arbea, JI & Jordana, R in Jordana, R & Ariño, AH, 1994:22) Weiner, WM, 1996:185
Species duprei (Beruete, E, Arbea, JI & Jordana, R in Jordana, R & Ariño, AH, 1994:26) Weiner, WM, 1996:186
Species maiteae (Beruete, E, Arbea, JI & Jordana, R in Jordana, R & Ariño, AH, 1994:28) Weiner, WM, 1996:186
Species minutus (Denis, 1932) Weiner, WM, 1996:185, t.t.
Species sorogainensis (Beruete, E, Arbea, JI & Jordana, R in Jordana, R & Ariño, AH, 1994:24) Weiner, WM, 1996:185

Genus Allonychiurus Yoshii, 1995
Species borensis (Beruete, E, Arbea, JI & Jordana, R in Jordana, R & Ariño, AH, 1994:30) Pomorski, RJ, 2002:il
Species caprariae (Dallai, 1969) Pomorski, RJ, 2002:il
Species donjiensis (Lee, B-H & Kim, J-T, 1994:208) Pomorski, RJ, 2002:il
Species flavescens (Kinoshita, 1916) Yoshii, 1995
Species foliatus (Rusek, J, 1967:189) Pomorski, RJ, 2002:il
Species hangchowensis (Stach, J, 1964:8) Pomorski, RJ, 2002:il
Species indicus (Choudhuri, 1965) Pomorski, RJ, 2002:il
Species jindoensis (Lee, B-H & Kim, J-T, 1994:211) Babenko, AB, 2007:369
Species jongaksanensis (Weiner, 1989) Weiner, WM, 1996:182
Species kimi (Lee, 1973) Yoshii, 1995
Species mariangeae (Thibaud & Lee, 1994) Pomorski, RJ, 2002:il
Species mediaseta (Lee, 1974) Weiner, WM, 1996:182
Species michelbacheri (Bagnall, 1948) Pomorski, RJ, 2002:il
Species pamirensis (Martynova, 1975) Pomorski, RJ, 2002:il
Species pseudocellitriadis (Lee, 1974) Weiner, WM, 1996:182
Species sensilatus (Thibaud & Massoud, 1980) Pomorski, RJ, 2002:il
Species sensitivus (Handschin, 1928) Pomorski, RJ, 2002:il
Species shanghaiensis (Rusek, 1971) Pomorski, RJ, 2002:il
Species shinbugensis (Lee, 1974) Weiner, WM, 1996:182
Species spinosus (Bagnall, 1949) Pomorski, RJ, 2002:il
Species tianshanicus (Martynova, 1971) Pomorski, RJ, 2002:il
Species vandeli (Cassagnau, P, 1960:405) Pomorski, RJ, 2002:il

Genus Uralaphorura Martynova, EF, 1978:63
Species sp.1 Babenko, AB, 2007:372
Species schilovi (Martynova, 1976) Martynova, 1978, t.t.
Species tunguzica Babenko, AB, 2007:369
Species varicellata Babenko, 2009
Species yanensis Babenko, 2009

Tribus Onychiurini Börner, 1906

Genus Similonychiurus Pomorski, RJ, 2007:49
Species voegtlini (Christiansen, K & Bellinger, P, 1980:446) Pomorski, RJ, 2007:50, t.t.
Similonychiurus voegtlini from the USA
After Pomorski RJ, 2007 Fig.9


Genus Onychiurus Gervais, 1841 key
Remark by ICZN. 2007:575: Placed on the Official List of Names in Zoology. Opinion 291.

Onychiurus sp. from Japan
2008 © Tsukiji, T.

Species sp.1 Absolon, K, 1900:191
Species sp.2 Leruth, R, 1939:198, B
Species sp.3 Palacios-Vargas, JG et Díaz, M, 1996:168
Species sp.4 Pomorski, RJ et Steinmann, D, 2004:20
Species sp.5 Kaprus, IJ, Pomorski, RJ, Skarzynski, D et Potapov, MB, 2005:893
Species sp.6 Elliott, WR, 2007:143
Species sp.7 Elliott, WR, 2007:143
Species aguzouensis Deharveng, 1978
Species ambulans (Linnæus, C, 1758:609) Gervais, 1841, t.t., B
Anurophorus ambulans from Europe
After Nicolet, H., 1847 Pl.6 Fig.14.
Anurophorus fimetarius from Switzerland
After Nicolet, H., 1842 Pl.5 Fig.2.
Lipura ambulans from the UK
After Lubbock J, 1873 Pl.43 Fig.1
Lipura ambulans from the UK
After Lubbock J, 1873 Pl.43 Fig.2
Podura terrestris nivea from Sweden
After de Geer, C., 1778 Tom.VII Pl.3 Fig.5.
Onychiurus ambulans from the UK
2002 © Hopkin, S.P.
Remark by Dányi, L. & Traser, G. 2008:23: All data published before description of Onychiurus circulans Gisin, 1952 are dubious.


  Subspecies inermis Ågren, 1903
  Subspecies intercedens Latzel, 1917,is
Species arans Gisin, 1952, B
Species ariegicus Deharveng, 1978
Species bhattii Yosii, 1963
Species boldorii Denis, 1937
Species cadaverinus (Handschin, 1921, i.s.) Handschin, 1921
Species certus (Nicolet, H, 1847:386) Giard, 1895
Remark by Ellis, W.N. & Bellinger, P.F. 1973:28: Nicolet, 1847 described new species from Chile. These are also described by Nicolet in Gay, 1851[lapsus for Nicolet in Gay, 1849]. But in the first paper Nicolet cites as author of the species Nicol. in Gay. Evidently the publications appeared in a reversed order.

Anurophorus certus from Chile
After Nicolet H in Gay C, 1854 Lam.1 Fig.4.

Species circulans Gisin, 1952, B
Remark by Dányi, L. & Traser, G. 2008:23: According to Pomorsky (1998) Onychiurus circulans is a junior synonym of Onychiurus ambulans (Linnaeus), but in our opinion it is a valid, separate taxon.

Species confugiens da Gama, 1962
Species darii Pomorski, 1998
Species decemsetosus Yosii, 1966
Species dissimulans Gisin, 1952, B
Species dubius (Nicolet, H, 1847:386) Giard, 1895
Remark by Ellis, W.N. & Bellinger, P.F. 1973:28: Nicolet, 1847 described new species from Chile. These are also described by Nicolet in Gay, 1851[lapsus for Nicolet in Gay, 1849]. But in the first paper Nicolet cites as author of the species Nicol. in Gay. Evidently the publications appeared in a reversed order.

Species fimetarius Denis, 1938
Remark by Gisin, H. 1960:140: Species inquirenda.

Species formosanus Denis, 1929
Species humboldtianus (Weiner, 1986) Pomorski, RJ, 2002:1
Species insinuans Gisin, 1952
Species jaceki Kaprus, IJ, 2009:60
Onychiurus jaceki from Kyrgyzstan
After Kaprus IJ, 2009 Fig.4A

Species justi Denis, 1938
  Subspecies porteri Tomlin, 1975
Species longipes von Olfers, 1907
Species movile Gruia, 1989
Species obsiones Cassagnau, 1963
Species orienteuropeus Kaprus, IJ, 2009:58
Onychiurus orienteuropeus from Ukraine
After Kaprus IJ, 2009 Fig.3A

Species ortus Denis, 1935
Species pancici Curcic, BPM et Lucic, LR, 1999:190, i.s.
Species paolettii Dallai, 1975
Species penetrans Gisin, 1952
Species polychaetosus (Lee & Park, 1986) Park, K-H & Lee, B-H, 1995:443
Species ravanicae Curcic, BPM et Lucic, LR, 1998:71, i.s.
Species rectospinatus Stach, 1922

Onychiurus wilchi from the USA
After Wray, D.L., 1950 Pl.5 Fig.18.
Species reluctus Christiansen, K, 1961:61

Species saphianus Nguyen, 2001
Species sarmaticus Kaprus, IJ, 2009:62
Onychiurus sarmaticus from Ukraine
After Kaprus IJ, 2009 Fig.5A

Species stillicidii (Schiödte, 1849) Latzel, 1922
Species subambulans Denis, 1935
Species subcirculans Gisin, H, 1962:17
Species sublegans Gisin, 1960, B
Species trojan Curcic, BPM et Lucic, LR, 1998:70, i.s.
Species vtorovi Martynova, 1976
Onychiurus vtorovi from Kyrgyzstan
After Kaprus IJ, 2009 Fig.1A


Genus Deuteraphorura Absolon, K, 1901:387
Species acicindelia (Salmon, JT, 1958:710) Pomorski, RJ, 2002:il
Species angelieri (Izarra, 1968) Pomorski, RJ in Christiansen, K, 2001:il
Species ameskoana (Beruete, E, Arbea, JI & Jordana, R, 2001:22) Pomorski, RJ, 2002:il
Species antheuili (Denis, 1936) Pomorski, RJ in Christiansen, K, 2001:il
  Subspecies aellenii Gisin, 1960
  Subspecies thesauri Gisin, 1960
Species apuanica (Dallai, R, 1970:446) Pomorski, RJ, 2002:il
Species aralarensis (Beruete, E, Arbea, JI & Jordana, R, 2001:17) Pomorski, RJ, 2002:il
Species arminiaria (Gisin, 1961) Christian, E, 2002:264
Species austriaria (Gisin, H, 1962:14) Christian, E, 2002:264
Species banatica (Gruia, 1965) Pomorski, RJ in Christiansen, K, 2001:il
Species bergamaria (Gisin, 1956) Pomorski, RJ in Christiansen, K, 2001:il
Species boneti (Gisin, 1953) Pomorski, RJ in Christiansen, K, 2001:il
Species bosnaria (Gisin, H, 1964:394) Pomorski, RJ in Christiansen, K, 2001:il
Species cebennaria (Gisin, 1956) Pomorski, RJ, 1996:52
Remark by Janssens, F 2000.04.30: "cebennaria" is presently depriciated. In this checklist, "inermis" is established as the tentatively valid name of "cebennaria", in line with the proposal of Bellinger (2000:in litt.) to resolve the invalid state of "Deuteraphorura".

Remark by Fjellberg, A. 2004.10.04:in litt.: But until a neotype for "inermis" is designated, there is no published evidence that "cebennaria" is a junior synonym and according to ICZN rulings, until the appropriate actions are taken, "cebennaria" should be used in taxonomic publications.

Species closanica (Gruia, 1965) Pomorski, RJ in Christiansen, K, 2001:il
Species defensaria (Gisin, H, 1964:397) Pomorski, RJ in Christiansen, K, 2001:il
Species diaelleni (Neuherz & Nosek, 1976) Pomorski, RJ, 2002:il
Species difficilis (Dallai, 1969) Pomorski, RJ in Christiansen, K, 2001:il
Species doneztebensis (Beruete, E, Arbea, JI & Jordana, R, 2001:19) Pomorski, RJ, 2002:il
Species eduardi (Denis, 1937) Pomorski, RJ in Christiansen, K, 2001:il

Onychiurus fazii from Lebanon
After Christiansen, K., 1956 Fig.4.
Species fazii (Christiansen, K, 1956:130) Pomorski, RJ in Christiansen, K, 2001:il

Species frasassii (Fanciulli, 1999) Pomorski, RJ in Christiansen, K, 2001:il
Species galani (Beruete, E, Arbea, JI & Jordana, R, 2001:16) Pomorski, RJ, 2002:il
Species gangjinensis (Lee, B-H & Kim, J-T, 1995:138) Pomorski, RJ, 2002:il
Species gemae (Simón-Benito, JC & Luciáñez, MJ in Jordana, R & Ariño, AH, 1994:61) Pomorski, RJ in Christiansen, K, 2001:il
Species ghidinii (Denis, 1938) Pomorski, RJ in Christiansen, K, 2001:il
Species gigoni (Gisin, H, 1962:6) Pomorski, RJ in Christiansen, K, 2001:il
Species gridellii (Denis, 1938) Pomorski, RJ in Christiansen, K, 2001:il
Species handschini (Denis, 1924) Pomorski, RJ in Christiansen, K, 2001:il
  Subspecies strinatii Gisin, 1963
Species harrobiensis (Beruete, E, Arbea, JI & Jordana, R, 2001:15) Pomorski, RJ, 2002:il
Species hategana (Gruia, M, 1971:283) Pomorski, RJ in Christiansen, K, 2001:il
Species haybachae (Gisin, H, 1962:8) Pomorski, RJ in Christiansen, K, 2001:il
Species hussoni (Denis, 1935) Pomorski, RJ in Christiansen, K, 2001:il
Species imperfecta (Denis, 1938) Pomorski, RJ in Christiansen, K, 2001:il, B
Species inermis (Tullberg, 1869) Absolon, 1901, t.t., B
Anurophorus fimetarius from Europe
After Nicolet,, 1847 Pl.6 Fig.15.
Lipura fimetaria from the UK
After Lubbock J, 1873 Pl.46
Onychiurus fimetarius from New Zealand
After Salmon, J.T., 1941 Pl.47 Fig.130.
Remark by Dányi, L. & Traser, G. 2008:21: Deuteraphorura fimetaria (Linnaeus, 1766) is a "species inquirenda according to Gisin (1960)". [Janssens, F., 2008.11.15. Contrary to what Dányi & Traser (2008:21) claim, Gisin (1960:140) refers to "fimetarius" sensu Denis (1938), not to fimetaria von Linné, 1767. The latter is considered as a species dubia by Gisin (1960:151)].

Deuteraphorura inermis from the UK
Post-antennal organ
2002 © Hopkin, S.P.
Deuteraphorura inermis from the UK
2006 © Valentine., B
Deuteraphorura inermis from the UK
2009.04.15© Robertson, A.
Deuteraphorura inermis from the UK
2009.04.15© Robertson, A.
Deuteraphorura inermis from France
2009.04.14© Raphael from Paris
Deuteraphorura inermis from the UK
2009.04.29© Robertson, A.

  Subspecies dentiferus Kos, 1939
Species inferni Gisin, 1956
Species insubraria (Gisin, 1952) Pomorski, RJ in Christiansen, K, 2001:il
Deuteraphorura insubraria
2006 © Boldsystems.org

Species jana (Christiansen, K & Bellinger, P, 1980:463) Pomorski, RJ in Christiansen, K, 2001:il
Species jitkae (Rusek, 1964) Pomorski, RJ in Christiansen, K, 2001:il
Species kosarovi (Zonev, 1973) Pomorski, RJ in Christiansen, K, 2001:il
Species kratochvili (Nosek, 1963) Pomorski, RJ in Christiansen, K, 2001:il
Species labainensis (Beruete, E, Arbea, JI & Jordana, R, 2001:21) Pomorski, RJ, 2002:il
Species leitzaensis (Beruete, E, Arbea, JI & Jordana, R, 2001:13) Pomorski, RJ, 2002:il
Species lusa (Christiansen, K & Bellinger, P, 1980:465) Pomorski, RJ in Christiansen, K, 2001:il
Species mangazeya Babenko, AB, 2007:372
Species meziadica (Gruia, 1972) Pomorski, RJ in Christiansen, K, 2001:il
Species mildneri (Christian, E, 1985:146) Pomorski, RJ in Christiansen, K, 2001:il
Species nervosa (Stach, J, 1954:167) Pomorski, RJ in Christiansen, K, 2001:il
Species nevoi (Gruia, Poliakov & Broza, 2000) Pomorski, RJ in Christiansen, K, 2001:il
Species opa (Christiansen, K & Bellinger, P, 1980:466) Pomorski, RJ in Christiansen, K, 2001:il
Species oregonensis (Denis, 1929) Pomorski, RJ in Christiansen, K, 2001:il
Species ossaria (Gisin, H, 1964:395) Pomorski, RJ in Christiansen, K, 2001:il
Species paro (Christiansen, K & Bellinger, P, 1980:468) Pomorski, RJ in Christiansen, K, 2001:il
Species pseudobosnaria (Dallai, R, 1970:447) Pomorski, RJ in Christiansen, K, 2001:il
Species pseudofimetaria (Folsom, 1917) Pomorski, RJ in Christiansen, K, 2001:il
  Varietas talpae Delamare Deboutteville C, 1947:239
Species pseudoghidinii (Dallai, 1969) Pomorski, RJ in Christiansen, K, 2001:il
Species pseudoinsubraria (Dallai, R, 1970:447) Pomorski, RJ in Christiansen, K, 2001:il
Species quadrisilvaria (Gisin, H, 1962:16) Christian, E, 2002:264
Species rendsinae (Haybach, 1962) Pomorski, RJ in Christiansen, K, 2001:il
Species romanica (Gruia, 1965) Pomorski, RJ in Christiansen, K, 2001:il
Species scotaria (Gisin, 1954) Pomorski, RJ, 1996:49
Deuteraphorura scotaria from the UK
Male genital plate
2002 © Hopkin, S.P.
Deuteraphorura scotaria from the UK
Female genital plate
2002 © Hopkin, S.P.

Species sexpunctata (Schäffer, 1897) Pomorski, RJ, 2002:il
Species silesiaca (Dunger, W, 1977:10) Pomorski, RJ, 1998:163
Species silvaria (Gisin, 1952) Weiner, WM, 1996:185
Species spipolae Massera, 1949
Species traiani Gruia & Popa, 2005
Species trisilvaria (Gisin, H, 1962:15) Christian, E, 2002:264
Species troglocarpathica Absolon, K, 1901:381
Remark by Absolon, K. 1901:381: "...welche ich später beschreiben werde."

Remark by Salmon, J.T. 1964:189: Species indeterminatae.

Remark by Janssens, F. 2009.01.29: Given Absolon did not provide a proper description of this Deuteraphorura species the name troglocarpathica is a nomen nudum.

Species variabilis (Stach, J, 1954:164) Pomorski, RJ, 1996:53
Species vercoraria (Gisin, 1963) Pomorski, RJ in Christiansen, K, 2001:il
Species vinuensis (Simón-Benito, JC & Luciáñez, MJ in Jordana, R & Ariño, AH, 1994:67) Pomorski, RJ in Christiansen, K, 2001:il
Species zalbidensis (Beruete, E, Arbea, JI & Jordana, R, 2001:20) Pomorski, RJ, 2002:il

Genus Orthonychiurus Stach, 1954
Species afghanistanensis (Stach, J, 1960:512) Pomorski, RJ in Christiansen, K, 2001:il
Species ancae (Gruia, M, 1971:285) Pomorski, RJ in Christiansen, K, 2001:il
Species argaensis (Arbea, JI & Jordana, R in Jordana, R & Ariño, AH, 1994:40) Pomorski, RJ in Christiansen, K, 2001:il
Species azoricus (Jacquemart, S, 1974:1) Pomorski, RJ, 2002:il
Species bunsteri (Loksa & Rubio, 1966) Pomorski, RJ in Christiansen, K, 2001:il
Species camerunensis (Schött, 1926) Pomorski, RJ in Christiansen, K, 2001:il
Species cunhai (Arlé, R, 1970:1) Pomorski, RJ, 2002:il
Species doinae (Gruia, 1972) Pomorski, RJ in Christiansen, K, 2001:il
Species durkeei (Christiansen, K & Bellinger, P, 1980:458) Pomorski, RJ in Christiansen, K, 2001:il
Species folsomi (Schäffer, 1900) Najt, J & Weiner, WM in Najt, J & Matile, L, 1997:18
Species gloriensis (Gruia, M, 1987:151) Pomorski, RJ in Christiansen, K, 2001:il
Species gurjakhanii Choudhuri, DK, 1958:150
Species himalayensis (Choudhuri, DK, 1958:147) Pomorski, RJ in Christiansen, K, 2001:il
Species izuruensis (Yosii, R, 1956:51) Pomorski, RJ in Christiansen, K, 2001:il
Species kowalskii (Stach, J, 1964:12) Pomorski, RJ in Christiansen, K, 2001:il
Species lampedusae (Fanciulli & Dallai, 1995) Pomorski, RJ in Christiansen, K, 2001:il
Species neocaledonicus Najt, J et Weiner, WM in Najt, J et Matile, L, 1997:18

Onychiurus novae-zealandiae(sic) from New Zealand
After Salmon, J.T. 1942 Pl.12 Fig.1.
Species novaezealandiae (Salmon, JT, 1942:158) Pomorski, RJ in Christiansen, K, 2001:il
Species pseudostachianus (Gisin, 1956) Pomorski, RJ, 1998:182
Remark by Pomorski, R.J. 1998:182: The separateness of Orthonychiurus pseudostachianus [from Orthonychiurus stachianus (Bagnall, 1939)] should be confirmed by a detailed examination.

Species rectopapillatus (Stach, 1933) Pomorski, RJ, 1996:56, t.t.
Species richardsi (Choudhuri, DK, 1958:150) Pomorski, RJ, 2002:il
Species saasveldensis (Weiner & Najt, 1991) Weiner, WM, 1996:187
Species schoenviszkyi (Loksa, I, 1967:292) Pomorski, RJ in Christiansen, K, 2001:il
Species selgae (Simón-Benito, JC & Luciáñez, MJ in Jordana, R & Ariño, AH, 1994:65) Pomorski, RJ in Christiansen, K, 2001:il
Species stachianus (Bagnall, 1939) Pomorski, RJ, 1998:179
Species subantarcticus (Salmon, JT, 1949:15) Pomorski, RJ in Christiansen, K, 2001:il
Species subcadaverinus (Denis, 1931) Pomorski, RJ, 2002:il
Species sublapidarius (Stach, 1963) Pomorski, RJ in Christiansen, K, 2001:il
Species ugandensis (Choudhuri, DK, 1958:152) Pomorski, RJ, 2002:il

Genus Bionychiurus Pomorski, RJ, 1996:92
Species normalis (Gisin, 1949) Pomorski, RJ, 1996:92, t.t., B
Species oblongatus (Lee & Park, 1986) Pomorski, RJ, 2002:il
Species yongyeonensis (Yosii, R, 1966:550) Pomorski, RJ, 2002:il

Genus Deharvengiurus Weiner, WM, 1996:183
Species argus (Denis, 1924) Weiner, WM, 1996:183, t.t.
Species denisi (Stach, J, 1934:191) Pomorski, RJ, 1998:190
Species lopezi (Deharveng, L & Gouze, A, 1984:395) Weiner, WM, 1996:184
Species microchaetosus (Loksa, 1959) Pomorski, RJ, 2002:il
Species orghidani (Gruia, M, 1967:103) Pomorski, RJ in Christiansen, K, 2001:il
Species pseudocantabricus (Gouze, 1983) Pomorski, RJ in Christiansen, K, 2001:il
Species severini (Willem, V, 1902:277) Pomorski, RJ in Christiansen, K, 2001:il, B
Species subargus (Deharveng, L & Gouze, A, 1984:395) Weiner, WM, 1996:184
Species xerophilus (Loksa, I, 1964:86) Pomorski, RJ in Christiansen, K, 2001:il

Genus Onychiuroides Bagnall, 1948
Species anellii (Denis, 1938) Weiner, WM, 1996:186
Species beroni (Gruia, 1972) Pomorski, 2006
Species bojani Pomorski, 2006
Species bureschi (Handschin, 1928) Pomorski, 2006
Species canzianus (Stach, J, 1934:200) Pomorski, 2006
Species cavernicolus (Stach, J, 1934:212) Weiner, WM, 1996:186
Species cerberus Pomorski, 2006
Species granulosus (Stach, J, 1930:285) Pomorski, RJ, 1996:62
Species hauseri (Dallai, 1975) Pomorski, RJ in Christiansen, K, 2001:il
Species hrabei (Rusek, 1963) Pomorski, 2006
Species igori Pomorski, 2006
Species longisetosus (Stach, J, 1954:145) Pomorski, 2006 si
Species multisetis (Gruia, M, 1971:287) Gruia, 2003
Species novus (Stach, J, 1960:511) Pomorski, RJ in Christiansen, K, 2001:il
Species occultus Pomorski, 2006
Species paoloi Pomorski, 2006
Species papillaeferus (Stach, J, 1946:20) Pomorski, 2006
Species paucituberculatus (Stach, J, 1930:285) Pomorski, 2006
Species peteri Pomorski, 2006
Species pirinensis Pomorski, 2006
Species postumicus (Bonet, 1931) Pomorski, RJ, 1996:95, t.t.
Species pseudogranulosus (Delamare Deboutteville & Gisin, 1951) Weiner, WM, 1996:186, B
Species quadripapillatus (Rusek, 1965) Pomorski, 2006
Species subgranulosus (da Gama, 1964) Pomorski, 2006
Species vornatscheri (Stach, J, 1946:23) Pomorski, 2006

Genus Absolonia Börner, 1901
Species giganteus (Absolon, K, 1901:85) Börner, 1901, t.t.

Genus Argonychiurus Bagnall, 1949
Species bertrandi (Denis, 1936) Pomorski, RJ in Christiansen, K, 2001:il
Species bogheani (Gruia, 1989) Pomorski, RJ in Christiansen, K, 2001:il
Species fistulosus (Gisin, 1956) Weiner, WM, 1996:183
Species lenticularius (Gisin, H, 1962:5) Pomorski, RJ in Christiansen, K, 2001:il
Species perforatus (Handschin, 1920) Pomorski, RJ, 1996:94
Species papulosus (Gisin, H, 1964:392) Pomorski, RJ in Christiansen, K, 2001:il

Genus Ongulonychiurus Thibaud & Massoud, 1986
Species colpus Thibaud & Massoud, 1986, t.t.

Genus Vibronychiurus Pomorski, 1998 key
Species aestimabilis Pomorski, RJ, 2006:674
Species archivari (Christiansen, K, 1956:128) Pomorski, RJ, 2006:673
Species caucasicus Pomorski, RJ, 2006:675
Species hermonicus (Gruia, Poliakov & Broza, 2000) Pomorski, RJ, 2006:676
Species vinolentus Pomorski, 1998 tt

Genus Pilonychiurus Pomorski, RJ, 2007:56
Species delhezi (Stomp, 1983) Pomorski, RJ, 2007:56, t.t.

Tribus Cribrochiurini Weiner, 1996

Genus Cribrochiurus Weiner, WM, 1996:189
Species cribrosus (Gisin, 1957) Weiner, WM, 1996:189, t.t.
Species subcribrosus (Gisin, 1957) Weiner, WM, 1996:190



References