http://www.collembola.org/taxa/proiinae.htm - Last updated on 2009.09.15 by Frans Janssens
Checklist of the Collembola: Proisotominae

Subfamilia Proisotominae Stach, 1947
Genus Number
of species
Appendisotoma21
Archisotoma25
Arlea7
Ballistura23
Bonetrura1
Burmisotoma1
Clavisotoma14
Cliforga1
Dimorphotoma2
Folsomia159
Folsomides62
Folsomina5
Gnathisotoma4
Gnathofolsomia2
Guthriella1
Mucrotoma1
Narynia3
Proisotoma77
Pseudofolsomia2
Scutisotoma29
Strenzketoma1
Subisotoma14
Villusisotoma2
Weberacantha5
Proisotominae462
Genera with number of species
Folsomia candida from the UK
2003 © Hopkin, S.P.
Folsomia candida from the UK
2005 © CEH.


Genus Villusisotoma Christiansen, KA et Nascimbene, P, 2006:334, -
Species brevis Christiansen, KA et Nascimbene, P, 2006:335, -, t.t.
Species longa Christiansen, KA et Nascimbene, P, 2006:336, -

Genus Burmisotoma Christiansen, KA et Nascimbene, P, 2006:340, -
Species lamellifera Christiansen, KA et Nascimbene, P, 2006:342, -, t.t.

Genus Gnathisotoma Cassagnau, P, 1957:393
Species bicolor Cassagnau, P, 1957:393, t.t.
Species deharvengi Najt, 1981
Species pallens Yoshii, 1990
Species patagonica Yoshii, 1990

Genus Folsomia Willem, 1902 key
Folsomia candida from the UK
2005 © CEH.
Remark by ICZN. 2007:322: Placed on the Official List of Names in Zoology. Opinion 1431.

Species abrupta Potapov & Deharveng, 2000
Species achaeta Bagnall, 1939
Species agrelli Gisin, 1944
Species albens Kaprus & Potapov, 1999
Species alpigena Stach, 1947
Species alpina Kseneman, 1936
Species altaica Martynova, 1974:18
Species altamontana Yosii, 1971
Species amazonae Winter, 1963
Species americana Stach, 1947
Species amplissima Potapov & Babenko, 2000
Species ancestor Potapov & Babenko, 2000
Species annamariae Winter, 1963
Species antricola Loksa, 1959
Species arena Potapov MB et Babenko A in Potapov, MB, 2006:16
Folsomia arena from Russia
After Potapov MB, 2006 Fig.19

Species asiatica Martynova, 1971
Species atropolaris Potapov & Babenko, 2000
Species baicalica Stebaeva & Potapov, 1998
Species baida Potapov, MB, 2006:12
Folsomia baida from Russia
After Potapov MB, 2006 Fig.2

Species baijali Prabhoo, 1971
Species bashkira Potapov MB et Kuznetsova N in Potapov, MB, 2006:14
Folsomia bashkira from Russia
After Potapov MB, 2006 Fig.4

Species bidentata Lee, 1977
Species binoculata (Wahlgren, 1899)
Species bisetosa Gisin, 1953
Species bisetosella Fjellberg, A, 2005:111
Species bogojevicae Dunger, W et Zivadinovic, J, 1989:3
Species borealis Potapov & Babenko, 2000
Species brevicauda Agrell, 1939
Species brevifurca (Bagnall, 1949) Goto, HE & Lawrence, PN in Kloet, GS & Hincks, WD, 1964:7
Remark by Potapov, M. 2001:186: As was shown by Lawrence (1973), the species was described from a single juvenile, so it became a species dubia.

Species brevisensilla Potapov & Babenko, 2000
Species britannica Stach, 1947
Species candida Willem, V, 1902:280, t.t., B
Remark by Potapov, M. 2001:188: According to the accurate study of types by Lawrence (1973), F. litsteri, F. distincta and F. kingi (described as three independent species) are the growth stages of this large-sized species.

Folsomia candida from the UK
2006.04.08 © Read, D.
Folsomia candida from Germany
2006.12.26 © Kohl, F.
Folsomia candida from France, juvenile
2007 © Raveton, M.
Remark by Dányi, L. & Traser, G. 2008:47: Data of Stach (1929) and Geyer & Mann (1940) were corrected by Loksa (1960:49).

Folsomia candida from France
After Tully, T., 2008 ©
First instar 0.35mm
Folsomia candida from Spain
After Baquero, E., 2009.05.22 ©
In laboratory culture

  Varietas indistincta Green, 1964
Species checae Lucianez & Simon, 2003
Species ciliata Babenko & Bulavintsev, 1993
Species clavipila Cassagnau, P, 1954:619
Species coeruleogrisea (Hammer, 1938) Fjellberg, A, 2007:35
Species continentalis Potapov & Dunger, 2000
Species cryptophila Potapov & Babenko, 2000
Species decaxiophthalma Ford, 1962
Species decemoculata Stach, J, 1946:29
Species decophthalma Steiner, 1958
Species decopsis Steiner, 1958
Species diplophthalma (Axelson, 1902) Axelson, 1907, B
  Varietas caeca Kseneman, 1935:12
  Varietas pallida Kseneman, 1938:462-463
  Varietas tetrophthalma Kseneman, 1935:12
  Forma anocellata Maynard EA, 1951:87
Species dovrensis Fjellberg, 1976
Species duodecimoculata Ford, 1962
Species echinosetosa Stebaeva, 1978
Species elongata (MacGillivray, 1896) James, 1933
Species evansi Fjellberg, 1984
Species fimetaria (Linnæus, C, 1758:609) Willem, 1925:626, B
Species fimetarioides (Axelson, WM, 1903:8) Handschin, 1929:59, B
Species gebhardti Loksa, I, 1964:95
Species giustii Dallai, R, 1970:451
Species hasegawai Yosii, R, 1959:117
Species heterocellata Stebaeva & Potapov, 1998
Species hexasetosa Lee, B-H, 1977:157
Species hidakana Uchida, H et Tamura, H, 1968:7
Species highlandia Wray, DL, 1950:95
Species hissarica Martynova, 1971
Species hodgei Maynard, EA, 1951:86
Species hoffi Scott, HG, 1961:264
Species hrabei Rusek, 1984
Species hubeiensis Ding, Huang & Chen, 2006
Species inoculata Stach, J, 1946:34
Folsomia inoculata from the UK
Post-antennal organ
2004 © Hopkin, S.P.
Folsomia inoculata from the UK
Basal dental ridges toothlike extended
2008.04.04 © Shaw, P.
Folsomia inoculata from the UK
2+2 anterior manubrial setae
2008.04.04 © Shaw, P.

Species kerni Gisin, 1948
Species kirgisica Martynova, 1973
Species ksenemani Stach, 1947
Species kuramotoi Yoshii, 1995
Species kurushica Potapov, MB, Abdurakhmanov, GM et Aitekova, MM, 2001:?
Species longidens Potapov & Babenko, 2000
Species lunata Salmon, JT, 1943:73
Species macrochaetosa Martynova, 1977
Species macroseta Ford, 1962
Species magadani Martynova, 1977
Species manolachei Bagnall, 1939, Bhabitus lateral
Remark by Dányi, L. & Traser, G. 2008:48: The [Hungarian] data of Folsomia quadrioculata (Tullberg) reported by Dunger (1975) included "Folsomia nana Gisin" [= Folsomia manolachei] as well.

Species martynovae Potapov, 2001
Folsomia duodecimoculata from Ukraine
After Kaprus IJ, 1997 Fig.1A

Species microchaeta Agrell, I, 1939:5
Species mikhaili Jordana & Baquero, 2008
Species minipunctata Zhao & Tamura, 1992
Species minuta Lee, B-H et Kim, J-T, 1994:211
Species miradentata Salmon, JT, 1943:74
Species monoculata (Bagnall, 1949) Goto & Lawrence in Kloet & Hincks, 1964:7
Species monophthalma Bagnall, 1939
Species montana Martynova, 1971
Species montigena Stach, J, 1946:32
Species nakajimai Yosii, R, 1959:116
Species nanjingensis Ding, Ji & Chen, 2006
Species nigromaculata Najt, 1981
Species nivalis (Packard, 1873) Scott, HG, 1961:265
Species obscurocellata Potapov & Cassagnau, 2000
Species ocellata Jordana, R, 1980:101
Species octoculata Handschin, 1925
Species orientalis Martynova, 1977
Species ozeana Yosii, 1954
Species palaearctica Potapov & Babenko, 2000
Species paoinflata Potapov MB et Stebaeva S in Potapov, MB, 2006:18
Species penicula Bagnall, 1939
Species picea Christiansen & Tucker, 1977:374
Species potapovi Babenko, AB, 2007:375
Species prima Mills, HB, 1931:3
Folsomia prima from the USA
2009.03.07 © Bergen, L.
Folsomia prima from the USA
2009.03.07 © Bergen, L.

Species pseudodecemoculata Stebaeva, 1971
Species pseudodiplophthalma Stach, 1947
Species pusilla Salmon, JT, 1944:143
Species pyrenaea Cassagnau, P, 1954:622

Species quadrioculata (Tullberg, 1871) Linnaniemi, 1907, B


Folsomia quadrioculata from New Zealand
After Salmon, J.T. 1941 Pl.49 Fig.154.
Folsomia quadrioculata from the UK
2005 © Brocklehurst, K.

Species regularis Hammer, 1953
Species riozoyoshiii Potapov & Cassagnau, 2000
Species rossica Potapov & Dunger, 2000
Species salmoni Stach, 1947
Species santokhi (Baijal, HN, 1958:351)
Bagnallela santokhi from the Himalaya
After Baijal HN, 1958 Fig.3A.

Species sedecimoculata Salmon, JT, 1943:75
Species sensibilis Kseneman, 1934
Species setifrontalis Potapov & Marusik, 2000
Species setosa Gisin, 1953
Species setula Christiansen & Tucker, 1977:376
Species sexoculata (Tullberg, 1871) Handschin, 1929:57, B
  Subspecies britannica Stach, 1947
Species similis Bagnall, 1939:57, B
Species sp. Christiansen, K, 1958:69
Species sp. Frati, F, Fanciulli, PP, Carapelli, A et Dallai, R, 1997:53
Species sp. González, V, Díaz, M et Palacios-Vargas, JG, 1997:46
Species sp. Brand, RH et Dunn, CP, 1998:238
Species sp. Potapov, MB, 2000:235
Species sp. Potapov, MB, 2000:235
Species sp. Potapov, MB, 2000:235
Species sp. Potapov, MB, 2000:235
Species sp. Potapov, MB, 2000:235
Species sp. Potapov, MB, 2000:235
Species sp. Potapov, MB, 2000:235
Species sp. Potapov, MB, 2000:236
Species sp. Potapov, MB, 2000:236
Species sp. Potapov, MB, 2000:236
Species sp. Potapov, MB, 2000:236
Species sp. Potapov, MB, 2000:236
Species sp. Potapov, MB, 2000:236
Species sp. Potapov, MB, 2000:236
Species sp. Potapov, MB, 2000:236
Species sp. Potapov, MB, 2000:236
Species sp. Potapov, MB, 2000:236
Species sparsosetosa Potapov & Stebaeva, 1997
Species spinosa Kseneman, 1936, B
Species stebaevae Potapov MB et Stevaeva S in Potapov, MB, 2006:19
Folsomia stebaevi(sic) from Russia
After Potapov MB, 2006 Fig.43

Species stella Christiansen & Tucker, 1977:379
Species strenzkei Nosek, 1963
Species sylvia Wray, DL, 1953:148
Species taigicola Stebaeva & Potapov, 1998
Species taimyrica Martynova & al., 1973
Species tatarica Martynova, 1964
Folsomia tatarica from Russia
After Potapov MB, 2006 Fig.3

Species tesari Dunger, 1970
Species thalassophila Bagnall, 1940
Species tianschanica Martynova, 1969
Species torpeda Potapov MB et Taskaeva A in Potapov, MB, 2006:11
Folsomia torpeda from Russia
After Potapov MB, 2006 Fig.1

Species trisetata Jordana, R et Ardanaz, A, 1981:41
Species ulrikeae Najt & Thibaud, 1987
Species variabilis Fjellberg, 1984
Species vasilyevi Berezina, OG et Potapov, MB, 2006:9
Species villosa Potapov & Marusik, 2000
Species volgensis Martynova, 1967
Species vtorovi Martynova, 1971
Species wanxianensis Ji, M, Ding, Y-F et Chen, J-X, 2007:59
Species wellingdae Potapov, M et Culik, MP, 2002:69

Genus Gnathofolsomia Deharveng, L et Christian, E, 1984:97
Species contrapunctata (Kseneman, 1950) Potapov, MB, 2000:236
Species palpata Deharveng, L et Christian, E, 1984:98, t.t.

Genus Pseudofolsomia Martynova, 1967
Species acanthella Martynova, 1967
Species spinata (Martynova, 1964)

Genus Arlea Womersley, H, 1939:319
Species adetolai de Mendonça, MC, Abrantes, EA et Fernandes, LH, 2006:58
Species arenicola Abrantes, EA et de Mendonça, MC, 2005:936
Species caeca Rapoport & Rubio, 1968
Species lucifuga (Arlé, 1939) Womersley, H, 1939:319, t.t.
Species psammophila de Mendonça, MC, Abrantes, EA et Fernandes, LH, 2006:57
Species spinisetis Mendonça & Arlé, 1987
Species tridens Barra, J-A, 1997:472

Genus Folsomina Denis, 1931 key
Species infelicia Greenslade, P, 1999:4
Folsomina infelicia from Malaysia
After Greenslade, P., 1999 Fig.1.

Species lawrencei Greenslade, P, 1999:6
Folsomina onychiurina from the Solomon Islands
After Lawrence, P.N., 1969 Fig.11.

Species onychiurina Denis, 1931, t.t., B
Folsomina yosiii from the Solomon Islands
After Lawrence, P.N., 1969 Fig.19.

Species wuyanensis Zhao & Tamura, 1992
Remark by Greenslade, P. 1999:12: The status of this species is left unchanged, although it is possible that F. wuyanensis is a synonym of F. onychiurina.

Species yongxingensis Chen, 1987

Genus Guthriella Börner, C, 1906:172
Species muskegis (Guthrie, JE, 1903:72) Börner, C, 1906:172, t.t.
Isotoma muskegis from the USA, ♂
After Guthrie JE, 1903 Plate XIII Fig.1
Isotoma muskegis from the USA, ♀
After Guthrie JE, 1903 Plate XIII Fig.2
Isotoma muskegis from the USA, ♀
After Guthrie JE, 1903 Plate XIII Fig.3
Guthriella muskegis from Canada
2009.07.17 Palacios-Vargas, J.G.


Genus Appendisotoma Stach, 1947
Species abiskoensis (Agrell, 1939)
Species absoloni (Rusek, 1966) Rusek, J in Dlabola, J, 1977:14
Species aera Christiansen & Bellinger, 1980
Species bisetosa Martynova, 1969
Species bulbosa (Folsom JW in Mills, HB, 1934:52) Nosek, 1964
Species dubia (Christiansen & Bellinger, 1981) Palacios-Vargas & Mejía, 1988:20
Species europaea von Törne, 1955
Species franzi (Haybach, 1962) Dányi, L & Traser, G, 2008:44
Species juliannae Traser, G, Thibaud, J-M et Najt, J, 1993:222
Species macgillivrayi (Dalla Torre, 1895)
Species mitra Uchida, H et Tamura, H, 1968:347
Species montana Martynova, 1969
Species nidicola (Agrell, 1939) Potapov, 2001
Species obsoleta (Mac Gillivray, 1896)
Species sibirica Stebaeva, 1985
Species tridentata (Baijal, HN, 1958:353) Potapov, 2001
Salmonia tridentata from the Himalaya
After Baijal HN, 1958 Fig.4A.

Species veca (Wray, 1952) Fjellberg, A, 1984:86

Appendisotoma vesiculata from the USA
2006 © Bernard, E.
Species vesiculata (Folsom, JW, 1937:57) Fjellberg, A, 1984:86, t.t.

Species woodi Christiansen & Bellinger, 1980
Species stebayevae (Grinbergs, A, 1962:63) Potapov, 2001
Species xinjiangica (Hao & Huang, 1995) Potapov, 2001

Genus Archisotoma Linnaniemi, WM, 1912:118 key
Species besselsi (Packard, AS, 1877:52) Linnaniemi, WM, 1912:119, t.t.
  Subspecies uniseriata Strenzke, 1955
Species bothrilongaequalis Thibaud & Ndiaye, 2006
Species brucei (Carpenter, GH, 1907:474) Womersley, H, 1936:321
Isotoma Brucei(sic) from Antarctica
After Carpenter GH, 1907 Plate I Fig.1
Archisotoma brucei from Antarctica
After Carpenter, 1907 in Greenslade, P., 2006 Fig.61.1.

Species catiae Abrantes, EA et de Mendonça, MC, 2007:55
Species gourbaultae Thibaud, 1993
Species heraultensis Christian, E et Thibaud, J-M, 1996:94
Species interstitialis Delamare Deboutteville, 1953
Species litoralis Thibaud, J-M et Weiner, WM in Najt, J et Matile, L, 1997:81
Species madagascariensis Thibaud, J-M, 2008:513
Species martae Fjellberg, A et Jucevica, E, 2000:21
Species megalops Bagnall, 1939
Species nigricans Bagnall, 1939
Species pauliani Delamare Deboutteville, 1953
Species poinsotae da Gama, 1968
Species polaris Fjellberg & Poinsot, 1975
Species pulchella (Moniez, 1890) Denis, 1923, B
Species quadrioculata Fjellberg, A, 1988:40
Archisotoma quadrioculata from Norway
After Fjellberg, A, 1988 Fig.4A

Species sabulosa Barra, J-A, 1997:474
Species senegalensis Thibaud & Ndiaye, 2006
Species subtheae Thibaud & Boumezzough, 2006
Species theae Fjellberg, 1980
Species utinomii Yosii, 1971
Species vaoensis Thibaud, J-M et Weiner, WM in Najt, J et Matile, L, 1997:81
Species vareli Sterzynska & Ehrnsberger, 2000
Species xavieri Cardoso, MA, 1966:84

Genus Cliforga Wray, 1952
Species alleghaniensis Wray, 1952

Genus Proisotoma Börner, 1901
Species sp.1 Potapov, M, Babenko, A et Fjellberg, A, 2006:74
Species sp.2 Potapov, 2001
Species alticola (Loksa & Rubio, 1966) Mari Mutt, JA & Bellinger, PF, 1990:64
Species andina Rapoport & Rubio, 1968
Species anopolitana Schulz, H-J et Lymberakis, P, 2006:230
Species atrata (Salmon, JT, 1941:314)
Species bayouensis Mills, HB, 1931:1
Species beta Christiansen & Bellinger, 1980
Species biseta Rapoport, 1963
Species brevidens Stach, J, 1947:204
Proisotoma brevidens from Poland
After Stach, J., 1947 Plate XX Fig.1.

Species brevipenna (Mac Gillivray, 1896)
Species bulba Christiansen & Bellinger, 1980
Species burti Fernando, 1960
Species caerulea Salmon, 1955
Species carolii (Tarsia, 1940)
Species clavipila (Axelson, WM, 1903:7) Linnaniemi, 1912
Species coeca da Gama, 1961
Species davidi Barra, J-A, 2001:23
Species decemoculata (Scherbakov, 1899) Denis, JR, 1931:118
Species dottrensi Gisin, 1952
Species erasoi Izarra, 1968
Species fernandeziana Rubio, 1974
Species fraterna Rusek, J, 1967:193
Species frisoni Folsom, JW, 1937:55
Species fungi Selga, 1962
Species granulata (Salmon, JT, 1943:379) Lawrence
Species granulata (Stach J, 1947:227) Stach, 1955
Proisotoma granulata from Poland
After Stach, J., 1947 Plate XXV Fig.1.

Species haweaensis (Salmon, JT, 1941:314)
Species himalayana Baijal, HN, 1958:353
Proisotoma himalayana from the Himalaya
After Baijal HN, 1958 Fig.5A.

Species immersa (Folsom, JW, 1924:1) Denis, JR, 1931:119
Isotoma immersa from the USA
After Folsom JW, 1924 Fig.1

Species intermedia Izarra, 1969
Species intermixta Handschin, 1928
Species juaniae Lucianez & Simón, 1992
Species judeana Palissa, 2006
Species koepckei Winter, 1967
Species lutii Izarra, 1973
Species macedoi Winter, 1967
Species mauretanica Handschin, 1925
Species micrura Börner, 1907
Species minima (Absolon, K, 1901:33) Linnaniemi, 1907, B
Proisotoma minima from Poland
After Stach, J., 1947 Plate XXI Fig.1.

Species minuta (Tullberg, 1871) Börner, 1903, t.t., B
Proisotoma minuta from Poland
After Stach, J., 1947 Plate XXII Fig.1.
Proisotoma minuta from the UK
2005 © Hopkin, S.P.
Proisotoma minuta from the UK
2005 © CEH.
Proisotoma minuta from the UK
1+1 anterior manubrial setae
6 anterior dental setae
2005 © Hopkin, S.P.
Proisotoma minuta from the USA
2007 © Murray, T.
Proisotoma minuta from the UK, juvenile
2007.10.03 © Valentine, B
Proisotoma minuta from the UK
2007.12.08 © Valentine., B
Proisotoma minuta from the UK
2007.12.28 © Valentine., B
Proisotoma minuta from the UK
2007.12.30 © Valentine., B
Proisotoma minuta from the UK
2008.07.03 © Valentine., B
Proisotoma minuta from the UK
2008.11.12 © Valentine., B
Proisotoma minuta from France
2008.12.07 © Jacquemin, G.
Proisotoma minuta juv. from the USA
2009.03.03 © Justis, S.
Proisotoma minuta juv. from the USA
2009.03.03 © Justis, S.

Species multituberculata Martynova, 1971
Species musci Lawrence, PN, 1969:547
Proisotoma musci from the Solomon Islands
After Lawrence, P.N., 1969 Fig.1.

Species muscicola Stach, J, 1965:353
Species najtae Selga, 1971
Species niger (Carpenter, GH, 1925:8)
Species nigromaculosa Folsom, JW, 1932:62
Species nova Palissa, 1969
Species octojuga Salmon, JT, 1949:33

Cryptopygus okukensis from New Zealand
After Salmon, J.T. 1941 Pl.48 Fig.141.
Species okukensis (Salmon, JT, 1941:313)
Species oliveirae Deharveng, L, 1984:123
Species pajonalica Loksa & Rubio, 1966
Species palermitana (Börner, 1903) Denis, JR, 1931:119
Species pallida (Moniez, 1894) Denis, JR, 1931:118, s.i.
Species papillosa Stach, 1937
Species paronai Börner, 1907
Species perparva Jackson, 1927
Species pettersonae Christiansen, KA et Nascimbene, P, 2006:340, -
Species quinocellata Salmon, 1954
Species ramosi Arlé, R, 1960:1960:166
Species rhopalotricha Massoud & Rapoport, 1968
Species ridleyi Denis, 1933
Species ripicola Linnaniemi, WM, 1912:128
Species roberti Fjellberg, 1991
Species sagardoyi Izarra, 1981, n.n.
Species santosorum Palacios-Vargas, JG et Arbea, JI, 2009:449
Proisotoma santosorum from Mexico
After Palacios-Vargas JG & Arbea JI Fig.9

Species senetijohani Baijal & Chandra, 1970
Species sepulcralis (Folsom, JW, 1902:364) Poole, RW in Poole, RW & Gentili, P, 1997:52
Species subdeflexa Delamare Deboutteville, 1951
Species subminuta Denis, JR, 1931:112
Proisotoma subminuta from Spain
After Arbea, J.I., 2006 Fig.1

Species submuscicola Nguyen, 1995
Species tenella (Reuter, 1895) Linnaniemi, 1912
Proisotoma tenella from Poland
After Stach, J., 1947 Plate XXVI Fig.1.

Species terrigena (Salmon, JT, 1943:379)
Species turikana Palacios-Vargas, JG et Arbea, JI, 2009:446
Proisotoma turikana from Venezuela
After Palacios-Vargas JG & Arbea JI Fig.1

Species vestita (Brown, 1923)
Species violenta Banasco, 1985,cf
Species xanthella Salmon, JT, 1949:34

Genus Folsomides Stach, 1922 key
Folsomides sp. from Australia
Scale bar = 0.1 mm
1999 © Walter, D.E.
Folsomides sp. from Australia
Head; scale bar = 0.01 mm
1999 © Walter, D.E.
Folsomides sp. from Indonesia
2008.06.29 © Nurtjahya, E.

Species almanzorensis Lucianez & Simón, 1991
Species alvarezi Selga, 1960
Species analuisae Fjellberg, A, 1993:470
Species andinensis Díaz, A et Najt, J, 1984:139
Species angularis (Axelson, WM, 1905:791) Gisin, 1942, B
Species arenosus Martynova, 1979
Species arenus Potapov & Thibaud, 2003
Species aridoviator Potapov & Stebaeva, 1997
Species arnoldi Suhardjono & Greenslade P, 1994:22
Species asiaticus Martynova, 1970
Species ayllonensis Simón & Luciáñez, 1990
Species cenkeh Yoshii, R et Suhardjono, YR, 1992:12
Species centralis (Denis, JR, 1931:111) Greenslade P & Thompson, 1981
Folsomides centralis from Vanuatu
2006 © Boldsystems.org

Species chichinautzini Kovác, L et Palacios-Vargas, JG, 1996:601
Species croci Potapov & Thibaud, 2003
Species cumbrosus Fjellberg, A, 1993:463
Species decemoculatus (Scherbakov, 1899) Ellis, WN, 1974:118
Species deflexus Schött, 1927
Species delamarei Thibaud, Najt & Jacquemart, 1994
Species denisi (Womersley, H, 1935:215) Greenslade, P in Houston, WWK, 1994:72
Astephanus denisi from Australia
After Womersley H, 1935 Fig.4e

Species deserticolus Wood, TG, 1970:79
Folsomides deserticola(sic) from Australia
After Wood, T.G, 1970 Fig.1

Species famarensis Fjellberg, A, 1993:467
Species graminis Fjellberg, A, 1993:466
Species insularis Thibaud, J-M et Weiner, WM in Najt, J et Matile, L, 1997:85
Species intermedius Fjellberg, A, 1993:468
Species lawrencei Gers & Deharveng, 1985
Species marchicus (Frenzel, 1941) Stach, 1947
Species mediterraneus Arbea, JI et Jordana, R, 2002:2
Species monosetis Massoud & Rapoport, 1968
Species mosambicensis Cardoso, MA, 1966:87
Species nanus Ellis, WN, 1974:115
Species neozealandia Salmon, 1948
Species nepalicus Yosii, 1971
Species nigrocellatus Fjellberg, A, 1993:466
Species ononicolus Fjellberg, A, 1993:467
Species oromii Fjellberg, A, 1993:466
Species parvulus Stach, 1922, t.t.
Folsomides parvus from Poland
After Stach, J., 1947 Plate X Fig.1.
Folsomides exiguus from Australia
After Womersley H, 1935 Fig.4b
Folsomides exiguus
2006 © Boldsystems.org
Folsomides parvulus
2006 © Boldsystems.org

Species petiti (Delamare Deboutteville, 1948) Wood, TG, 1970:81
Species pinicolus Fjellberg, A, 1993:468
Species pocosensillatus Fjellberg, A, 1993:465
Species portucalensis da Gama, 1961
Species pseudangularis Chen, 1985
Species quinocellatus Salmon, 1954
Species semiparvulus Fjellberg, A, 1993:460
Species sexophthalmus (Womersley, 1934) Wood, TG, 1970:82
Species socorrensis Kovác, L et Palacios-Vargas, JG, 1996:598
Species sp. Greenslade, P in Houston, WWK, 1994:72
Species sp. Arbea, JI et Blasco-Zumeta, J, 2001:41
Species supranubius Fjellberg, A, 1993:469
Species teno Fjellberg, A, 1993:466
Species teres Christiansen & Bellinger, 1980
Species terrus Fjellberg, A, 1993:464
Species tonellus Fjellberg, A, 1993:468
Species troglobius (Rapoport & Maño, 1969) Palacios-Vargas, JG, 1999:36
Species unicus Fjellberg, A, 1993:464
Species urumqiensis Hao & Huang, 1995
Species vinosus Fjellberg, A, 1993:462
Species viridescens Pomorski, RJ, 2001:265
Species virungiensis Martynova, 1978
Species xerophilus Fjellberg, A, 1993:462
Species yucatanicus Kovác, L et Palacios-Vargas, JG, 1996:595
Species zairensis Martynova, 1978

Genus Scutisotoma Bagnall, 1949 key
Species acorrelata Potapov, M, Babenko, A et Fjellberg, A, 2006:41
Species ananevae (Babenko & Bulavintsev, 1992) Potapov, M, Babenko, A & Fjellberg, A, 2006:16
Species armeriae (Fjellberg, 1976) Potapov, M, Babenko, A & Fjellberg, A, 2006:16
Species baica Potapov, M, Babenko, A et Fjellberg, A, 2006:49
Species christianseni (Stach, 1959) Potapov, M, Babenko, A & Fjellberg, A, 2006:16
Species dodecocellata Potapov, M, Babenko, A et Fjellberg, A, 2006:42
Species fjellbergi (Dunger, WG, 1982:54) Potapov, M, Babenko, A & Fjellberg, A, 2006:16
Species indigirka Potapov, M, Babenko, A et Fjellberg, A, 2006:51
Species karadagi Potapov, M, Babenko, A et Fjellberg, A, 2006:24
Species kolymica Potapov, M, Babenko, A et Fjellberg, A, 2006:28
Species ladaki (Denis, 1936) Potapov, M, Babenko, A & Fjellberg, A, 2006:21
Species longisensilla Potapov, M, Babenko, A et Fjellberg, A, 2006:30
Species matalini Potapov, M, Babenko, A et Fjellberg, A, 2006:36

Scutisotoma millimetrica from Russia
After Potapov, M. et al. 2006 Fig.80.
Species millimetrica Potapov, M, Babenko, A et Fjellberg, A, 2006:31
Species montana Potapov, M, Babenko, A et Fjellberg, A, 2006:45
Species muriphila (Grinbergs, A, 1968:139) Potapov, M, Babenko, A & Fjellberg, A, 2006:19
Species nicksmeti Potapov, M, Babenko, A et Fjellberg, A, 2006:52
Species oirota (Vilkamaa, 1988) Potapov, M, Babenko, A & Fjellberg, A, 2006:16
Species potapovi Xie, G-L et Chen, J-X, 2008:?
Species schisti Potapov, M, Babenko, A et Fjellberg, A, 2006:34
Species sp.1 Potapov, M, Babenko, A et Fjellberg, A, 2006:74
Species stachanoremi Potapov, M, Babenko, A et Fjellberg, A, 2006:47
Species stepposa (Martynova, 1975) Potapov, M, Babenko, A & Fjellberg, A, 2006:16
Species subarctica (Gisin, 1950) Potapov, M, Babenko, A & Fjellberg, A, 2006:16
Species tenuidentifera Potapov, M, Babenko, A et Fjellberg, A, 2006:39
Species tianshanica (Martynova, 1971) Potapov, M, Babenko, A & Fjellberg, A, 2006:17
Species titusi (Folsom, JW, 1937:49) Potapov, M, Babenko, A & Fjellberg, A, 2006:17, t.t.
Species tolerans (Babenko, 1994) Potapov, M, Babenko, A & Fjellberg, A, 2006:17
Species variabilis (Gisin, 1949) Potapov, M, Babenko, A & Fjellberg, A, 2006:17
  Subspecies psammophila Loksa, I et Bogojevic,J, 1970:128

Genus Mucrotoma Rapoport & Rubio, 1963
Species leuca Rapoport & Rubio, 1963, t.t.

Genus Bonetrura Christiansen & Bellinger, 1980
Species boneti (Yosii, 1962) Palacios-Vargas, 1983, t.t.

Genus Subisotoma Stach, 1947
Species asiatica (Martynova, 1971) Fjellberg, 1993
Species australis (Womersley, 1934)
Species corsica Poinsot & Barra, 1982
Species loftyensis (Womersley, 1934)
Species meridionalis (Dallai, R, 1973:510)
Species navacerradensis (Selga, 1962)
Species pratensis (Zivadinovic, 1972)
Species pusilla (Schäffer, 1900) Stach, J, 1947:111, t.t.
Subisotoma pusilla from Poland
After Stach, J., 1947 Plate XII Fig.1.

Species sp.3 Potapov, M, Babenko, A et Fjellberg, A, 2006:74
Species sp.4 Potapov, M, Babenko, A et Fjellberg, A, 2006:74
Species sp.5 Potapov, M, Babenko, A et Fjellberg, A, 2006:74
Species sp.6 Potapov, M, Babenko, A et Fjellberg, A, 2006:74
Species tenuis (Dunger, WG, 1982:51) Fjellberg, 1993
Species veletensis (Steiner, 1959) Potapov, 2001

Genus Ballistura Börner, C, 1906:172
Ballistura sp. from Estonia
2008.05.14 © Tartes, U.

Species alpa Christiansen & Bellinger, 1980
Species albertinae (Ellis, 1976:291) Potapov, 2001
Species antiqua Folsom, JW, 1937:34

Proisotoma aqualata from New Zealand
After Salmon, J.T. 1941 Pl.53 Fig.220.
Species aqualata (Salmon, JT, 1941:333) Salmon, JT, 1945:69

Species bengalensis Yosii, 1966
Species borealis (Axelson, WM, 1905:791) Potapov, 2001
Species ewingi (James, 1933) Ellis, 1970
Species excavata (Folsom, JW, 1937:41)
Species extra (Christiansen & Bellinger, 1980) Fjellberg, A, 1984:86
Species fitchi (Denis, 1933) Potapov, 2001
Species gisini (da Gama, 1964) Potapov, M, Babenko, A & Fjellberg, A, 2006:13
Species hankoi (Stach, J, 1930:288) Dányi, L & Traser, G, 2008:45
Species libra (Christiansen & Bellinger, 1980) Skidmore, 1995:54
Species nilgiris (Denis, 1937) Potapov, 2001
Species obtusicauda (Schäffer, 1897)
Species pratensis (Betsch, J-M et Massoud, Z, 1970:210)
Species schoetti (von Dalla Torre, 1895) Stach, J, 1947:237, t.t.
Isotoma litoralis(sic) from Sweden
After Schött H, 1893 Taf.VII Fig.1
Ballistura schoetti from Sweden
2006 © Hall, K.
Ballistura schoetti from Sweden
2006 © Hall, K.
Ballistura schoetti from France
2008.06.25 © Lebeaux, P.

  Subspecies lutea Womersley, 1934
Species sp. Dunger, WG, 1982:57
Species sp. González, V, Díaz, M et Palacios-Vargas, JG, 1997:47
Species tuberculata (Stach, J, 1947:218) Skidmore, 1995:55
Proisotoma tuberculata from Poland
After Stach, J., 1947 Plate XXIII Fig.1.

Species vernoga Wray, 1958
Species vetusta Folsom, JW, 1937:35
Species wrangeliensis Martynova, 1973

Genus Weberacantha Christiansen, KA, 1951:24 key
Species beckeri (Stebaeva, 1966) Potapov, M, Babenko, A & Fjellberg, A, 2006:58

Weberacantha echinodermata from Russia
After Potapov, M. et al. 2006 Fig.168.
Species echinodermata Potapov, M, Babenko, A et Fjellberg, A, 2006:58

Species janetscheki (Yosii, 1971) Potapov, M, Babenko, A & Fjellberg, A, 2006:58

Weberacantha magnomucrella from Mongolia
After Potapov, M. et al. 2006 Fig.176.
Species magnomucrella Potapov, M, Babenko, A et Fjellberg, A, 2006:61

Weberacantha octa from Alaska
After Christiansen, K., 1951 Fig.1.
Species octa Christiansen, KA, 1951:25, t.t.

Genus Strenzketoma Potapov, M, Babenko, A et Fjellberg, A, 2006:63
Species buddenbrocki (Strenzke, 1954) Potapov, M, Babenko, A & Fjellberg, A, 2006:63, t.t.

Genus Dimorphotoma Grinbergs, A, 1975:101
Species huadongensis (Chen, 1985) Potapov, 2001
Species porcella Ellis, WN, 1976:295

Genus Narynia Martynova, 1968
Species andreevae Martynova, 1975
Species kolymiensis Martynova, 1981
Species setosa Martynova, 1968

Genus Clavisotoma Ellis, 1970
Species africana (Womersley, 1934)
Species brisbanensis (Womersley, H, 1935:215)
Species fatonei (Rapoport, 1959)

Clavisotoma filifera from the UK
2005 © CEH
Species filifera (Denis, JR, 1931:112)

Species fitchioides (Denis, 1947)
Species inaequalis (Schäffer, C, 1898:403)
Species lamelligera (Börner, C, 1909:108)
Species laticauda (Folsom, JW, 1937:42)
Species palustris (Cassagnau, P, 1959:82)
Species plicicauda (Handschin, 1927)
Species richardi (Denis, 1924)
Species sjoestedti (Wahlgren, 1910)
Species yamorensis (Uchida, 1949)
Species yosii (Stach, 1947)



References