http://www.collembola.org/taxa/tomoceri.htm - Last updated on 2009.09.14 by Frans Janssens
Checklist of the Collembola: Tomoceridae

Familia Tomoceridae Schäffer, 1896 key
Genus Number
of species
Antennacyrtus1
Aphaenomurus1
Entomocerus1
Lasofinius2
Lepidophorella10
Lethemurus3
Monodontocerus3
Neophorella1
Novacerus3
Plutomurus25
Pogonognathellus8
Pseudolepidophorella1
Tomocerina12
Tomocerus70
Tomolonus1
Tritomurus2
Tomoceridae144
Genera with number of species
Pogonognathellus sp. from the UK
Body covered with iridescent scales
2007.12.19 © Campbell, A.
 
Tomocerus minor from the UK
3rd abdominal segment superequal to 4th
2008.09.24 © Robertson, A.


Subfamilia Tomocerinae Schäffer, 1896
Tomocerinae from Japan
2005.09.07 © Keresztes, G.
Tomocerus sp. from France
Long 3rd antennal segment
2007.02.23 © Ollivier, E.
Tomocerinae juv. from Belgium
2008.06.09 © Vuijlsteke, M.
 
Tomocerinae juv. from the UK
First instar with short 3rd antennomere
2008.09.28 © Barton, T.
Tomocerinae juv. from the USA
Long 3rd antennal segment
2009.03.14 © Justis, S.
Tomocerinae juv. from the UK
4th antennal segment > 1st + 2nd antennal segment
2009.04.20 © Robertson, A.


Tribus Tomocerini Salmon JT, 1941:346

Genus Tomocerus Nicolet, H, 1842:67
Remark by ICZN. 2007:821: Placed on the Official List of Names in Zoology. Opinion 239.

Tomocerus minor from the UK
Long 3rd antennal segment subcylindrical
2008.09.24 © Robertson, A.
Tomocerus minor from the UK
1st and 2nd antennal segments scaled
2008.12.22 © Valentine., B
Tomocerus minor from the UK
Long 3rd antennal segment subcylindrical
2009.01.26 © Robertson, A.
Tomocerus minor from Germany
Colour of 3rd antennal segment distinctly different from other segments
2009.01.26 © Peters, J.
Tomocerus sp. from the USA
Eyecluster with 6 visible ocelli
2009.02.26 © Cowen, R.

Species asahinai Yosii, 1954
Species asiaticus Martynova, 1969
Species asoka Yosii & Ashraf, 1965
Species baicalensis Martynova, 1969
Species baudoti Denis, 1932
Species bimaculatus Sun, Liang & Huang, 2006
Species caputiviolaceus Lee, 1975
Species catalanus Denis, 1924
Species changbaishanensis Wang, 1999
Species cheni Ma, Y-T et Christiansen, KA, 1998:47
Species christianseni Stomp, N, 1969:695
Species cucullatus Olfers, 1907, t.t.
Species cuspidatus Börner, C, 1909:114
Species denticulus Lee, 1975
Species deongyuensis Lee, 1975
Species emeicus Liu, Hou & Li, 1999
Species folsomi Denis, 1929
Species fopingensis Sun & Liang, 2008
Species hexipunctatus Sun, Liang & Huang, 2007
Species ishibashii Yosii, 1954
Species jesonicus Yosii, 1967
Species jezoensis Matsumura & Ishida in Matsumura, 1931:200
Species jordanai Sun, Liang & Huang, 2006
Species kinoshitai Yosii, 1954
  Subspecies dentiferus Yosii, R, 1956:90
Species lamelliger (Börner, 1903) Gisin, 1960:254
Species laxalamellus Lee, 1975
Species lividus Tullberg, T, 1877:32
Species maculatus Sun, Liang & Huang, 2007
Species magadanicus Martynova, 1979
Species magnus Sun & Liang, 2008
Species maximus Liu, Hou & Li, 1999
Species minor (Lubbock, 1862) Schäffer, 1900, t.t., B
Tomocerus plumbeus from the UK
After Lubbock, J., 1873 Pl.19.
Tomocerus plumbeus from the UK
After Beck J in Lubbock J, 1873 Pl.68
Tomocerus minor from Belgium
Tridentate dental spines
Legit Leruth, R., 1933.07.16
2008.05.28 © Janssens, F.
Tomocerus minor from Belgium
Unscaled 3rd antennal segment
Legit Leruth, R, 1933.10.03
2008.05.28 © Janssens, F.
Tomocerus minor from Belgium
2004 © Vuijlsteke, M.
Tomocerus minor from the UK
2005 © Hopkin, S.P.
Tomocerus minor from the UK
2006 © Cornwall, N.J.
Tomocerus minor from Belgium
2006 © Vuijlsteke, M.
Tomocerus minor from Germany
2006 © Peters, J.
Tomocerus minor from the UK
2007 © Barton, T.
Tomocerus minor from Germany
2007.02.15 © Peters, J.
Tomocerus minor from the USA
2007.05.04 © Moorehead, C.
Tomocerus minor from Belgium
2007.07.07 © Vuijlsteke, M.
Tomocerus minor from the UK
2007.07.22 © Valentine., B
Tomocerus minor from the USA
2007.09.21 © Moorehead, C.
Tomocerus minor from the UK
2007.12.14 © Botting, J.
Tomocerus minor from the Netherlands
2008.01.24 © Baas, A.H.
Tomocerus minor from the UK
2008.02.03 © Birch, P.
Tomocerus minor from the Netherlands
2008.02.03 © Baas, A.H.
Tomocerus minor from the UK
2008.02.27 © Spears, G.
Tomocerus minor from Belgium
2008.03.15 © Oger, P.
Tomocerus minor from the UK
2008.03.17 © Robertson, A.
Tomocerus minor from Belgium
2008.03.22 © Vuijlsteke, M.
Tomocerus minor from Belgium
2008.03.22 © Vuijlsteke, M.
Tomocerus minor from Belgium
2008.03.24 © Vuijlsteke, M.
Tomocerus minor from the USA
2008.05.29 © Nanz, S.
Tomocerus minor from the USA
2008.09.13 © Robertson, A.
Tomocerus minor from the UK
2008.09.24 © Robertson, A.
Tomocerus minor juvenile from the UK
2008.09.24 © Robertson, A.
Tomocerus minor from the UK
2008.10.21 © Spears, G.
Tomocerus minor from the UK
2009.01.14 © Valentine., B
Tomocerus minor from Germany
2009.02.18 © Peters, J.
Tomocerus minor from the UK
2009.02.27 © Knight., B
Tomocerus minor from Belgium
2009.03.15 © Janssens, F.
Tomocerus minor from the UK
2009.04.29 © Robertson, A.

Species mitrai Prabhoo & Muraleedharan, 1980
Species montanus (Oudemans, 1890:74) Yayuk, 1989:9
Species monticolus Huang & Yin, 1981
Species multisetus Sun, Liang & Huang, 2007
Species nepalicus Yosii, 1971
Species nodentalis Nguyen, 1995
Species noricus Latzel, 1918
Species obscurus Huang & Yin, 1981
Species ocreatus Denis, 1948
Species orcinus Absolon, K, 1901:88
Species orientalis Martynova, 1979
Species parvus Huang & Yin, 1981
Species petalospinus Salmon, 1969
Species problematicus Cassagnau, P, 1964:741
Species punctatus Yosii, 1967
Species purpuratus Sun, Liang & Huang, 2006
Species sachalinensis Martynova, 1979
Species serratospinus Salmon, 1969
Species setoserratus Salmon, JT, 1941:347
Species shuense Liu, 2003
Species sibiricus Reuter, 1891
Species similis Chen, J-X et Ma, Y-T, 1997:157
Species sp. I da Gama, 1959
Species sp. II da Gama, 1959
Species spinistriatus Lee, 1975
Species spinulus Chen, J-X et Christiansen, KA, 1998:51
Species steinbocki Yosii, 1971
Species taeniatus (Koch & Berendt, 1854 -) -
Species teldanicus Gruia, 1995
Species terrestralis (Stach, 1922) Cassagnau, P, 1958:121
Species varius Folsom, 1899
Species vasconius Bonet, 1928
Species violaceus Yosii, R, 1956:91
Species viridescens Wankel, 1860
Species viridis Yosii, 1967
Species vulgaris (Tullberg, 1871) Brook, 1883, B
Tomocerus vulgaris from Germany
2005 © Menzel, E.
Tomocerus vulgaris from Germany
2005 © Kohl, F.
Tomocerus vulgaris from Germany
2005 © Peters, J.
Tomocerus vulgaris from the Netherlands
2007 © Hautus, M.A.
Tomocerus vulgaris from Belgium
2007.07.12 © Janssens, F.
Tomocerus vulgaris from Germany
2007.04.07 © Kohl, F.
Tomocerus vulgaris from the Netherlands
2007.09.18 © Arp, X.
Tomocerus vulgaris from Germany
2008.01.25 © Peters, J.
Tomocerus vulgaris from the UK
2008.01.27 © Barton, T.
Tomocerus vulgaris from Belgium
2008.08.28 © Janssens, F.
Tomocerus vulgaris from the Netherlands
2008.10.06 © Baas, A.H.
Tomocerus vulgaris from the Netherlands
2008.10.07 © Baas, A.H.
Tomocerus vulgaris from the Netherlands
2008.10.08 © Baas, A.H.
Tomocerus vulgaris from the Netherlands
2008.10.10 © Baas, A.H.
Tomocerus vulgaris from France
2008.11.03 © Ramel, A.
Tomocerus vulgaris from Germany
2008.12.07 © Peters, J.
Tomocerus vulgaris from Belgium
2009.02.21 © Vuijlsteke, M.
Tomocerus vulgaris from the UK
2009.03.04 © Ransom, T.
Tomocerus vulgaris from Belgium
2009.03.07 © Janssens, F.
Tomocerus vulgaris from France
2009.04.19 © Lefebvre, V.
Det. Ponge, J.-F.
Tomocerus vulgaris from the USA
2009.09.01 © Hodge, K.

Species wushanensis Sun, Liang & Huang, 2007
Species zayueensis Huang & Yin, 1981

Genus Tritomurus Frauenfeld, 1854 key
Species falcifer Cassagnau, P, 1958:121
Species scutellatus Frauenfeld, 1854:15, t.t.

Genus Pogonognathellus Paclt, 1944
Pogonognathellus sp. from France
Furca with dental array
2007.04.10 © Lebeaux, P.
Pogonognathellus flavescens
from the Netherlands
Tapering antenna
2008.10.24 © Baas, A.H.
Pogonognathellus flavescens
from the Netherlands
2008.11.03 © Baas, A.H.

Species beckeri (Börner, C, 1909:115)
Pogonognathellus beckeri from Japan
2008 © Tsukiji, T.

Species bidentatus (Folsom, JW, 1913:463) Massoud, Z & Ellis, WN, 1974:292
Species borealis Yosii, 1967
Species celsus (Christiansen, 1964:670) Massoud, Z & Ellis, WN, 1974:293
Species dubius Christiansen, 1964:671
Pogonognathellus dubius from the USA
2005 © DiTerlizzi, T.
Pogonognathellus dubius from the USA
2005 © Murray, T.
Pogonognathellus dubius from the USA
2006 © Cresswell, S.
Pogonognathellus dubius from the USA
2007 © Murray, T.
Pogonognathellus dubius from Canada
2007.12.17 © McCann, S.
Pogonognathellus dubius from the USA
2008.09.09 © Bradford, A.
Pogonognathellus dubius from the USA
2008.10.08 © Bradford, A.
Pogonognathellus dubius from the USA
2008.12.06 © Houston, S.
Pogonognathellus dubius from the USA
2009.06.18 © Bradford, A.

Species elongatus Maynard, EA, 1951:126
Pogonognathellus elongatus from the USA
2006 © Cresswell, S.
Pogonognathellus elongatus from the USA
Body scales removed
2008.08.12 © Trently, D.

Species flavescens (Tullberg, 1871) Stach, 1929, B
Podura plumbea from Ireland
After Templeton, R, 1836 Pl.XI Fig.4,4.1
Pogonognathellus flavescens from Belgium
2004 © Vuijlsteke, M.
Pogonognathellus flavescens from the USA
2006 © Murray, T.
Pogonognathellus flavescens from the UK
2006.02.17 © Valentine., B
Pogonognathellus flavescens from Belgium
2006.03.19 © Vuijlsteke, M.
Pogonognathellus flavescens from the USA
2007.03.03 © Murray, T.
Pogonognathellus flavescens from the UK
2007.02.13 © Jones, D.
Pogonognathellus flavescens from the UK
2007.02.18 © Barton, T.
Pogonognathellus flavescens from the USA
2007.03.03 © Murray, T.
Pogonognathellus flavescens from the USA
2007.03.11 © Murray, T.
Pogonognathellus flavescens from Czechia
2007.06.23 © Krásenský, P.
Pogonognathellus flavescens from Czechia
2007.07.22 © Krásenský, P.
Pogonognathellus flavescens from the Netherlands
2007.11.05 © Baas, A.H.
Pogonognathellus flavescens from the UK
2007.11.14 © Campbell, A.
 
Pogonognathellus flavescens from the UK
2007.11.18 © Campbell, A.
 
Pogonognathellus flavescens from the UK
2007.11.25 © Warner, P.
Pogonognathellus flavescens from the Netherlands
2007.12.08 © Heemskerk, R.
Pogonognathellus flavescens from the Netherlands
2008.02.10 © Baas, A.H.
Pogonognathellus flavescens from the UK
2008.02.10 © Campbell, A.
Pogonognathellus flavescens from France
2008.02.11 © Lavoué., B
Pogonognathellus flavescens from the UK
Juvenile, descaled
2008.03.31 © Williams., B
Pogonognathellus flavescens from the UK
2008.05.01 © Farrell, S.
Pogonognathellus flavescens from Belgium
Scaled 3rd antennal segment
Legit Leruth, R, 1933.09.30
2008.05.28 © Janssens, F.
Pogonognathellus flavescens juv.
from the Netherlands
2008.10.15 © Baas, A.H.
 
Pogonognathellus flavescens juv.
from the Netherlands
2008.10.15 © Baas, A.H.
Pogonognathellus flavescens
from the Netherlands
2008.11.03 © Baas, A.H.
Pogonognathellus flavescens from France
2008.11.08 © Jacquemin, G.
Pogonognathellus flavescens from the UK
2009.01.31 © Ransom, T.
Pogonognathellus flavescens from Belgium
2009.02.21 © Vuijlsteke, M.

Species longicornis (Müller OF, 1776) Paclt, 1944, B
Tomocerus longicornis from the UK
After Beck J in Lubbock J, 1873 Pl.67
Tomocerus longicornis from the UK
After Lubbock J, 1873 Pl.17
Tomocerus plombeus(sic) from Europe
After Nicolet, H, 1847 Pl.6 Fig.12.
Pogonognathellus longicornis from the UK
2002 © Hopkin, S.P.
Pogonognathellus longicornis
2003 © Pettersson., B
Pogonognathellus longicornis from Sweden
2005 © Hall, K.
Pogonognathellus longicornis from the UK
2006 © Cornwall, N.J.
Pogonognathellus longicornis from the UK
2006 © Cornwall, N.J.
Pogonognathellus longicornis from Russia
2006 © Macroclub.ru.
Pogonognathellus longicornis from Germany
2006.09.30 © Kohl, F.
Pogonognathellus longicornis from Germany
2007 © Peters, J.
Pogonognathellus longicornis from Belgium
2007.02.07 © Lindsley, J.
Pogonognathellus longicornis from France
2007.06.09 © Lebeaux, P.
Pogonognathellus longicornis from Belgium
2007.09.08 © Oger, P.
Pogonognathellus longicornis from the UK
2007.10.13 © Brian.
Pogonognathellus longicornis from the UK
2007.10.13 © Brian.
Pogonognathellus longicornis from the UK
2007.11.10 © Campbell, A.
Pogonognathellus longicornis from Belgium
2007.11.24 © Carlet, G.
Pogonognathellus longicornis from Sweden
2007.12.04 © Hallmén, J.
Pogonognathellus longicornis from the Netherlands
2007.12.08 © Heemskerk, R.
Pogonognathellus longicornis from the Netherlands
2007.12.08 © Heemskerk, R.
Pogonognathellus longicornis from the UK
2008.04.03 © Spears, G.
Pogonognathellus longicornis from the UK
2008.07.27 © Michaelson, J.
Pogonognathellus longicornis from the UK
2008.09.01 © Talbot, M.
Pogonognathellus longicornis from the UK
2009.02.17 © Ransom, T.
Pogonognathellus longicornis from the UK
Unguis (U) with lateral teeth (LT)
Unguicular filament (uf) longer than unguis
2009.04.03 © Robertson, A.


Genus Tomolonus Mills, 1949

Tomolonus reductus from the USA
2006 © Murray, T.
Species reductus Mills, 1949, t.t.


Genus Monodontocerus Yosii, 1955
Species leqingensis Sun & Liang, 2009
Species modificatus (Yosii, 1955) Yosii, R, 1956:88, t.t.
  Subspecies satsumensis Yosii, R, 1956:89
Species odongnyeoensis Park, K-H et Lee, B-H, 1995:437

Genus Tomocerina Yosii, 1955 key
Species aokii (Yosii, R, 1972:94)
Species calcea Liu, Hou & Li, 1999
Species curta (Christiansen, 1964:662)
Species lamellifera (Mills, HB, 1934:84)
Species liliputana Yosii, 1967
Species minuta (Tullberg, T, 1877:32) Yosii, R, 1956:91
Species purpurithora Liu, Hou & Li, 1999
Species simplex Yosii, 1966
Species teres (Christiansen, K, 1964:653)
Species tianshanensis Ma, Y-T, Chen, J-X et Christiansen, K, 2003:512
Species tridentatus Sun, Liang & Huang, 2007
Species yiliensis Ma, Y-T, Chen, J-X et Christiansen, K, 2004:43

Genus Aphaenomurus Yosii, R, 1956:86
Species interpositus Yosii, R, 1956:86, t.t.
  Subspecies denticulatus Yosii, R, 1956:87

Genus Plutomurus Yosii, R, 1956:78
Species sp.1 Pomorski, RJ et Steinmann, D, 2004:20
Species sp.2 Kaprus, IJ, Pomorski, RJ, Skarzynski, D et Potapov, MB, 2005:894
Species abchasicus Martynova, 1969
Species baschkiricus (Skorikov, 1900)
Species belozerovi Martynova, 1979
Species borealis Suma, Y, 1981:502
Species brevimucronatus (Denis, 1928)
Species californicus (Folsom, JW, 1913:469)
Species carpaticus Rusek, J et Weiner, WM, 1978:742
Species diversispinus Yosii, R, 1966:555
Species edaphicus Yosii, 1967
Species grahami Christiansen, 1980:121
Species gul Yosii, R, 1966:552
Species iwatensis Yoshii, 1991
Species jeleznovozski Kniss & Thibaud, 1999
Species kelasuricus Martynova, 1969
Species leei Yosii, R, 1966:554
Species marmorarius Yosii, 1967
Species riugadoensis (Yosii, 1939) Yosii, R, 1956:78, t.t.
Species sorosi Kniss & Thibaud, 1999
Species suzukaensis (Yosii, 1938) Yosii, R, 1956:81
  Subspecies ehimensis Yosii, R, 1956:83
  Subspecies naikaiensis Yosii, R, 1956:83
  Subspecies ohminensis Yosii, R, 1956:82
Species unidentatus (Börner, 1901) Dobat, 1979, B
Species vigintiferispina Lee, 1974
Species wilkeyi Christiansen, K, 1964:659
Species yamatensis Yosii, R, 1956:84
  Subspecies kawasawai Yosii, R, 1956:85
  Forma albida Yosii R, 1956:85
  Subspecies kyushuensis Yosii, R, 1956:85

Genus Lethemurus Yosii, 1970
Species sp.1 Pomorski, RJ et Steinmann, D, 2004:20
Species finitimus Yosii, 1970
Species missus (Mills, 1949), t.t.

Genus Entomocerus Christiansen, KA et Pike, E, 2002:170, -
Species mirus Christiansen, KA et Pike, E, 2002:170, -, t.t.

Subfamilia Lepidophorellinae Absolon K, 1903

Tribus Lepidophorellini Womersley, 1939

Genus Lepidophorella Schäffer, 1897
Species australis Carpenter, GH, 1925:11
Species brachycephala (Moniez, 1894) Denis, 1923

Species communis Salmon, JT, 1937:353


Lepidophorella communis from New Zealand
After Salmon, J.T. 1937 Pl.49 Fig.d.
Lepidophorella communis from New Zealand
After Salmon, J.T. 1941 Pl.54 Fig.246.

Species flavescens (Nicolet, H, 1847:383) Schäffer, 1897
Cyphoderus giganteus from Chile
After Nicolet H in Gay C, 1854 Lam.1 Fig.7.
Remark by Ellis, W.N. & Bellinger, P.F. 1973:27-28: Cyphoderus flaveus Nicolet, 1847 is an incorrect original spelling (art.32a(ii)). Nicolet, 1847:383 described the new species Cyphoderus giganteus and C. flaveus. These are also described by Nicolet in Gay, 1851[lapsus for Nicolet in Gay, 1849]:92-93. In the last paper however "flavescens" is used in stead of "flaveus". But in the first paper Nicolet cites as author of both species Nicol. in Gay, evidently the publications appeared in a reversed order. This, and the fact that "flaveus" is bad latin, brings us to the conclusion that "flaveus" is an incorrect original spelling.


Species fusca Salmon, JT, 1941:340

Lepidophorella nigra from New Zealand
After Salmon, J.T. 1943 Pl.42 Fig.48.
Species nigra Salmon, JT, 1943:383
Species rubicunda Salmon, JT, 1941:342
Species spadica Salmon, JT, 1944:152
Species tiegsi (Womersley, H, 1942:26)
Millsia tiegsi from Australia
After Womersley, H., 1942 Fig.3A
Remark by Greenslade, P. in Dallai, R. 1989:110: The type series consists of three specimens, all probably early instars. Millsia is similar to Lepidophorella and differs only in the lack of scales and the shorter, more cylindrical dens. It is possible that Millsia is a synonym of Lepidophorella.

Species unadentata Salmon, JT, 1941:341

Genus Pseudolepidophorella Salmon, JT, 1941:342

Species longiterga (Salmon, JT, 1937:354) Salmon, JT, 1941:343, t.t.


Lepidophorella longiterga from New Zealand
After Salmon, J.T. 1937 Pl.49 Fig.h.
Pseudolepidophorella longiterga from New Zealand
After Salmon, J.T. 1941 Pl.54 Fig.259.


Genus Antennacyrtus Salmon, JT, 1941:343

Antennacyrtus insolitus from New Zealand
After Salmon, J.T. 1941 Pl.55 Fig.261.
Species insolitus Salmon, JT, 1941:344, t.t.


Genus Neophorella Womersley, 1934
Species dubia Womersley, 1934, t.t.

Genus Lasofinius Ireson & Greenslade P, 1990
Species gemini Ireson & Greenslade P, 1990:211
Species willi Ireson & Greenslade P, 1990:209, t.t.

Tribus Novacerini Salmon JT, 1941:345

Genus Novacerus Salmon, JT, 1942:259
Remark by Salmon, J.T. 1942:259: Novacerus Salmon JT, 1942 nomen novum to replace the name Neocerus Salmon, 1941, pre-occupied by Wasmann, 1893.

Species insolitatus (Salmon, JT, 1941:346) Salmon, JT, 1942:259

Neocerus spinosus from New Zealand
After Salmon, J.T. 1941 Pl.55 Fig.269.
Species spinosus (Salmon, JT, 1941:345) Salmon, JT, 1942:259, t.t.

Species tasmanicus (Womersley, H, 1937:154) Yosii, 1967
Tomocerus tasmanicus from Tasmania
After Womersley H, 1937 Fig.1g




References