Phylogenetics of Order Hymenoptera

reference

Sawflies, Bees, Ants, and Wasps

     =========================== Xyelidae 
     |
     |                    ====== Cimbicidae 
     |                    |
     |              ======|===== Diprionidae 
     |              |     |
     |              |     ====== Tenthredinidae 
     |              |
     |           ===|=========== Pterygophoridae  (= Perlidae) 
     |           |  |
     |           |  |=========== Argidae 
<<===|        ===|  |
     |        |  |  ============ Electrotomidae * 
     |  ======|  |
     |  |     |  =============== Blasticotomidae 
     |  |     |
     |  |     ================== Xyelotomidae * 
     |  |
     |  |     ================== Pamphiliidae 
     |  |     |
     |  |     |================= Megalodontidae 
     |  |=====|
     |  |     |================= Praesiricidae * 
     ===|     |
        |     ================== Xyelydidae * 
        |
        |     ================== Cephidae 
        |=====|
        |     ================== Sepulcidae * 
        |
        |        =============== Anaxyelidae 
        |     ===|
        |     |  =============== Gigasiricidae * 
        ======|
              |  =============== Siricidae 
              |  |
              ===|  ============ Xiphydriidae 
                 |  |
                 ===|     ====== Orussidae 
                    |  ===|
                    ===|  ====== Paroryssidae * 
                       |
                       ========= Apocrita 

* Extinction

Apocrita

     ============================== Tenthredinoidea (suborder)    
     |           
     |     ======================== Trigonalyoidea  
<<===|     |     
     |     |   ==================== Evanioidea
     ======|   |  
           |   |           ======== Cynipoidea   
           |   |        ===|
           |   |        |  ======== Proctotrupoidea
           ====|  ======| 
               |  |     |  ======== Chalcidoidea           
               |  |     ===|    
               |  |        ======== Platygasteroidea   
               ===|    
                  |      ========== Ichneumonoidea    
                  =======|    
                         ========== Aculeata

Aculeata

       ========= Chrysidoidea (wasps)    (suborder)
       |
<<=====|     === Sphecoidea --- Sphecidae (solitary wasps)
       |  ===|
       ===|  === Apoidea (bees)
          |
          ====== Vespoidea (wasps, ants)

Suborder Chrysidoidea (wasps)

              ====== Sclerogibbidae (family)
           ===|
           |  |  === Embolemidae 
           |  ===|
        ===|     === Dryinidae 
        |  |
        |  |  ====== Bethylidae 
     ===|  ===|
     |  |     ====== Chrysididae 
<<===|  |
     |  ============ Scolebythidae 
     |
     =============== Plumariidae 

Suborder Apoidea

     ======================== Stenotritidae (family)
<<===|   
     |======================= Colletidae   
     |    
     |           ============ Andrenidae    
     |           |
     |     ======|=========== Oxaeidae  
     |     |     |
     |     |     ============ Halictidae (sweat bees)
     ======|      
           |         ======== Melittidae   
           |   ======|    
           |   |     ======== Ctenoplectridae
           ====|    
               |     ======== Fideliidae           
               |  ===|    
               |  |  ======== Megachilidae (leaf cutting bees)   
               ===|    
                  |      ==== Anthophoridae (digger bees)    
                  =======|    
                         ==== Apidae 

Family Apidae (Bees)    

     =========== Euglossini (orchid bees) solitary (genus)
     |
<<===|    ====== Bombini (bumble bees)
     |    |
     ==x==|  === Meliponini (stingless bees)
          ===|
             === Apini (honey bees)
x = eusocial          

Suborder Vespoidea

           ============ Sapygidae (family) 
        ===|
     ===|  ============ Mutillidae 
     |  |
     |  =============== Tiphiidae 
     |
     |  =============== Sierolomorphidae 
     |  |
     |  |     ========= Rhopalosomatidae 
<<===|  |  ===|
     |  |  |  ========= Pompilidae (spider wasp)
     |  |  |
     ===|  |  ========= Bradynobaenidae 
        ===|  |
           |  |     === Scoliidae 
           ===|  ===|
              ===|  === Vespidae (hornets, yellow jackets)
                 |
                 ====== Formicidae (ants)

Family Formicidae (Ants)

                          ====== Aenictinae (genus)
                          |
                       ===|===== Aenictogitoninae 
                       |  |
                    ===|  ====== Dorylinae 
                    |  |
                 ===|  ========= Ecitoninae 
                 |  |
           ======|  ============ Cerapachyinae 
           |     |
           |     =============== Leptanilloidinae 
           |
        ===|==================== Apomyrminae 
        |  |
        |  |==================== Leptanillinae 
        |  |
        |  ===================== Ponerinae 
        |
     ===|     ================== Aneuretinae 
     |  |     |
     |  |  ===|================= Dolichoderinae 
     |  |  |  |
     |  |  |  ================== Formicinae 
     |  |  |
     |  ===|        ============ Myrmeciinae 
<<===|     |     ===|
     |     |  ===|  ============ Nothomyrmeciinae 
     |     |  |  |
     |     ===|  =============== Pseudomyrmecinae 
     |        |
     |        ================== Myrmicinae 
     |
     =========================== Formiciinae 

Family Vespidae (Wasps)

     =================== Euparagiinae (subfamily)
     |
<<===|  ================ Masarinae <pollen wasps> 
     ===|
        |  ============= Eumeninae <potter wasps>   
        ===|
           |      ====== Stenogastrinae <hover wasps> 
           ===x===| 
                  |  === Polistinae <paper wasps> 
                  ===|
                     === Vespinae <hornets & yellowjackets> 

x = social vespids