Vascular Plants
Ferns Life
Cycle
Sori form on the fronds. Each contains many sporangia mounted on stalks. Within each sporangium, the spore mother cells undergo meiosis producing four haploid spores each. When the humidity drops, The thin-walled lip cells of each sporangium separate. The annulus slowly straightens out. Then the annulus snaps forward expelling the spores. If a spore is blown to a suitable moist location, It germinates into a filament of cells. This grows into a prothallus with rhizoids, which absorb water and minerals from the soil; archegonia, which produce a single egg (by mitosis); antheridia, which form swimming sperm (again, by mitosis). If moisture is plentiful, the sperm swim to archegonia - usually on another prothallus because the two kinds of sex organs generally do not mature at the same time on a single prothallus.Fertilization restores the diploid number and begins a new sporophyte generation.The embryo sporophyte develops a foot that penetrates the tissue of the prothallus and enables the sporophyte to secure nourishment until it becomes self-sufficient. Although it is tiny, the haploid fern prothallus is a fully-independent, autotrophic plant.
The photo shows the sori on the underside of the leaflets of the Christmas fern.