Most biologists agree that animals evolved from simpler single-celled organisms. Exactly how this happened is unclear, because few fossils have been left to record the sequence of events. Faced with this lack of fossil evidence, researchers have attempted to piece together animal origins by examining the single-celled organisms alive today. Modern single-celled organisms are classified into two kingdoms: the prokaryotes and protists. Prokaryotes, which include bacteria, are very simple organisms, and lack many of the features seen in animal cells. Protists, on the other hand, are more complex, and their cells contain all the specialized structures, or organelles, found in the cells of animals. One protist group, the choanoflagellates or collar flagellates, contains organisms that bear a striking resemblance to cells that are found in sponges. Most choanoflagellates live on their own, but significantly, some form permanent groups or colonies. This tendency to form colonies is widely believed to have been an important stepping stone on the path to animal life. The next step in evolution would have involved a transition from colonies of independent cells to colonies containing specialized cells that were dependent on each other for survival. Once this development had occurred, such colonies would have effectively become single organisms. Increasing specialization among groups of cells could then have created tissues, triggering the long and complex evolution of animal bodies. This conjectural sequence of events probably occurred along several parallel paths. One path led to the sponges, which retain a collection of primitive features that sets them apart from all animals. Another path led to two major subdivisions of the animal kingdom: the protostomes, which include arthropods, annelid worms, mollusks, and cnidarians; and the deuterostomes, which include echinoderms and chordates. Protostomes and deuterostomes differ fundamentally in the way they develop as embryos, strongly suggesting that they split from each other a long time ago. Animal life first appeared perhaps a billion years ago, but for a long time after this, the fossil record remains almost blank. Fossils exist that seem to show burrows and other indirect evidence for animal life, but the first direct evidence of animals themselves appears about 650 million years ago, toward the end of the Precambrian period. At this time, the animal kingdom stood on the threshold of a great explosion in diversity. By the end of the Cambrian Period, 150 million years later, all of the main types of animal life existing today had become established. Moving onto Land When the first animals evolved, dry land was probably devoid of any kind of life, except possibly bacteria. Without terrestrial plants, land-based animals would have had nothing to eat. But when plants took up life on land over 400 million years ago, that situation changed, and animals evolved that could make use of this new source of food. The first land animals included primitive wingless insects and probably a range of soft-bodied invertebrates that have not left fossil remains. The first vertebrates to move onto land were the amphibians, which appeared about 370 million years ago. For all animals, life on land involved meeting some major challenges. Foremost among these were the need to conserve water and the need to extract oxygen from the air. Another problem concerned the effects of gravity. Water buoys up living things, but air, which is 750 times less dense than water, generates almost no buoyancy at all. To function effectively on land, animals needed support. In soft-bodied land animals such as earthworms, this support is provided by a hydrostatic skeleton, which works by internal pressure. The animal's body fluids press out against its skin, giving the animal its shape. In insects and other arthropods, support is provided by the exoskeleton (external skeleton), while in vertebrates it is provided by bones. Exoskeletons can play a double role by helping animals to conserve water, but they have one important disadvantage: unlike an internal bony skeleton, their weight increases very rapidly as they get bigger, eventually making them too heavy to move. This explains why insects have all remained relatively small, while some vertebrates have reached very large sizes. Speciation and Extinction Like other living things, animals evolve by adapting to and exploiting their surroundings. In the billion-year history of animal life, this process has created vast numbers of new species, each capable of using resources in a slightly different way. Some of these species are alive today, but these are a minority; an even greater number are extinct, having lost the struggle for survival. Speciation, the birth of new species, usually occurs when a group of living things becomes isolated from others of their kind (see Species and Speciation). Once this has occurred, the members of the group follow their own evolutionary path and adapt in ways that make them increasingly distinct. After a long period-typically thousands of years-their unique features mean that they can no longer breed with their former relatives. At this point, a new species comes into being. In animals, this isolation can come about in several different ways. The simplest form, geographical isolation, occurs when members of an original species become separated by a physical barrier. One example of such a barrier is the open sea, which isolates animals that have been accidentally stranded on remote islands. As the new arrivals adapt to their adopted home, they become more and more distinct from their mainland relatives. Sometimes the result is a burst of adaptive radiation, which produces a number of different species. In the Hawaiian Islands, for example, 22 species of honeycreepers have evolved from a single pioneering species of finch-like bird. Another type of isolation is thought to occur where there is no physical separation. In this case, differences in behavior, such as mate selection, may sometimes help to split a single species into distinct groups. If the differences persist for a long enough time, new species are created. The fate of a new species depends very much on the environment in which it evolved. If the environment is stable and no new competitors appear on the scene, an animal species may change very little in hundreds of thousands of years. But if the environment changes rapidly and competitors arrive from outside, the struggle for survival is much more intense. In these conditions, either a species changes, or it eventually becomes extinct. During the history of animal life, on at least five occasions, sudden environmental change has triggered simultaneous extinction on a massive scale. One of these mass extinctions occurred at the end of the Cretaceous Period, about 65 million years ago, killing all dinosaurs and perhaps two-thirds of marine species. An even greater mass extinction took place at the end of the Permian Period, about 200 million years ago. Many biologists believe that we are at present living in a sixth period of mass extinction, this time triggered by human beings.
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